NEWSLETTER 11/2014 03.11.2014

Please acknowledge use of the database www.shark-references.com in your publications, and cite: 

Pollerspöck, J. 2014, Bibliography database of living/fossil sharks, rays and chimaeras (Chondrichtyes: Elasmobranchii, Holocephali), www.shark-references.com, World Wide Web electronic publication, Version 2014


Pollerspöck J, Beaury B. 2014. Parasquatina zitteli nov. sp. (Elasmobranchii: Orectolobiformes) from the Maastrichian of Upper Bavaria (Gerhartsreiter Ditch, Siegsdorf) and remarks on the distribution of sharks of the order Orectolobiformes.Zitteliana, A54: 147-164

The genus Parasquatina is documented for the first time from the Gerhartsreiter Ditch near Siegsdorf (Upper Bavaria, Germany). The fossils are interpreted as a new species, for which the name Parasquatina zitteli nov. sp. is proposed, because they differ in morphology from all other species in that genus, especially with regard to the labial apron, which is absent in P. zitteli. Assignment of the genus Parasquatina to the order Squatiniformes is reversed, and the genus now placed in the order Orectolobiformes. In addition, a survey of
recent records of Parasquatina is given, together with an up-to-date overview of the stratigraphic distribution of the order Orectolobiformes, and resulting questions are discussed.

Diversity of Carpet sharks (Orectolobiformes) based on the recorded genera; x-axis: 
number of recorded fossil genera, y-axis: time


Plate: Stratigraphic Distribution of the Order Orectolobiformes.
Extinct genera are labeled with a cross (†) in front of the genus name. After the name of the genus you will find the number of valid species (extant and extinct). A continuous line documents fossil records of this genus. Broken line means missing fossil records.
Used abbreviation:
Hol = Holocene; Ple = Pleistocene; Pia = Piacenzian; Zan = Zanclean; Mes = Messinian; Tor = Tortonian; Srv = Serravallian; Lan = Langhian; Bur = Burdigalian; Aqt = Aquitanian; Cht = Chattian; Rup = Rupelian; Prb = Priabonian; Brt = Bartonian; Lut = Lutetian; Ypr = Ypresian; Tha = Thanetian; Sel = Selandian; Dan = Danian; Maa = Maastrichtian; Cmp = Campanian; San = Santonian; Con = Coniacian; Tur = Turonian; Cen = Cenomanian; Alp = Albian; Apt = Aptian; Brm = Barremian; Hau = Hauterivian; Vlg = Valanginian; Ber =
Berriasian; Tth = Tithonian; Kim = Kimmeridgian; Oxf = Oxfordian; Clv = Callovian; Bth = Bathonian; Baj = Bajocian; Aal = Aalenian; Toa = Toarcian; Plb = Pliensbachian; Sin = Sinemurian; Het = Hettangian;
Used references:
1 = Cappetta H. 2012. Handbook of Paleoichthyology, Vol. 3E: Chondrichthyes • Mesozoic
and Cenozoic Elasmobranchii: Teeth. Verlag Dr. Friedrich Pfeil, 512 S.
2 = Underwood CJ, Ward DJ, King C, Antar SM, Zalmout IS, Gingerich PD. 2011. Shark
and ray faunas in the Middle and Late Eocene of the Fayum Area, Egypt. Proceedings of
the Geologists' Association 122, 47-66.
3 = Guinot G, Cappetta H, Adnet S. 2014. A rare elasmobranch assemblage from the
Valanginian (Lower Cretaceous) of southern France. Cretaceous Research 48, 54-84.
4 = Adolfssen J, Ward DJ. 2014. Crossing the boundary: an elasmobranch fauna from
Stevns Klint, Denmark. Palaeontology, Palaeontology 57, 591-629.
5 = Guinot G, Underwood CJ, Cappetta H, Ward DJ. 2013. Sharks (Elasmobranchii:
Euselachii) from the Late Cretaceous of France and the UK. Journal of Systematic
Palaeontology 11, 589-671.
6 = Hamm SA, Cicimurri DJ. 2011. Early Coniacian (Late Cretaceous) selachian fauna from
the basal Atco Formation, lower Austin Group, north central Texas. Paludicola 8, 107-127.
7 = Kirkland JI, Eaton JG, Brinkman DB. 2013. Elasmobranchs from upper Cretaceous
freshwater facies in southern Utah. In: At the top of the Grand Staircase - The Late
Cretaceous of southern Utah. Loewen A, Titus L, Mark A. (eds.), Indian Univ. Press., 153-
194, fig. 9.1-9.22.
8 = Adolfssen J, Ward DJ. 2013. Neoselachians from the Danian (Early Paleocene) of
Denmark. Acta Palaeontologica Polonica, accepted manuscript,
9 = Canevet J-M. 2011. Les Chondrichthyens du Miocène moyen (Serravallien) de Salles
(Gironde, France). Cossmanniana 13, 59-79.
10 = Pimiento C, González-Barba G, Ehret DJ, Hendy AJW, Macfadden BJ, Jaramillo C.
2013. Sharks and Rays (Chondrichthyes, Elasmobranchii) from the Late Miocene Gatun
Formation of Panama. Journal of Paleontology 87, 755-774.
11 = Schultz O. 2013. Catalogus Fossilium Austriae Band 3 - Pisces Verlag der
Österreichischen Akademie der Wissenschaften, Wien, ISBN13: 978-3-7001-7238-3, 576
12 = Carrillo-Briceño JD, Aguilera OA, Rodriguez F. 2014. Fossil Chondrichthyes from the
central eastern Pacific Ocean and their paleoceanographic significance. Journal of South
American Earth Sciences 51, 76-90.
13 = Cook TD, Newbrey MG, Brinkman DB, Kirkland JI. 2014. Euselachians from the
freshwater deposits of the Hell Creek Formation of Montana. Geological Society of
America, Special Paper 503, 229-246, fig. 1-6, A1-A2, tabl. 1.
14 = Thies D, Müller A. 1993. A neoselachian fauna (Vertebrata, Pisces) from the Late
Cretaceous (Campanian) of Höver, near Hannover (NW Germany). Paläontologische
Zeitschrift 67, 89-107, 21 fig.
15 = Iserbyt A, De Schutter PJ. 2012. Quantitative analysis of Elasmobranch assemblages
from two successive Ypresian (Early Eocene) facies at Marke, western Belgium. Geologica
Belgica 15, 146-153.
16 = Bourdon J, Wright K, Lucas SG, Spielmann JA, Pence R. 2011. Selachians from the
Upper Cretaceous (Santonian) Hosta Tongue of the Point Lookout Sandstone, central New
Mexico. New Mexico Museum of Natural History and Science, Bulletin 52, 1-52.
17 = Jansen KR, Shimada K, Kirkland JI. 2013. Fossil Fish Fauna from the Uppermost
Graneros Shale (Upper Cretaceous: Middle Cenomanian) in Southeastern Nebraska.
Transactions of the Kansas Academy of Science 115, 145-152.
18 = Rees J. 2010. Neoselachian sharks from the Callovian-Oxfordian (Jurassic) of
Ogrodzieniec, Zawiercie Region, southern Poland. Palaeontology 53, 887-902.
19 = Delsate D, Weis R. 2010. La Couche à Crassum (Toarcien moyen) au Luxembourg:
stratigraphie et faunes de la coupe de Dudelange-Zoufftgen. In: Weis R, Guérin-Franiatte
S. (éditeurs) 2010. - Le Jurassique inférieur et moyen au Luxembourg - Nouvelles données
paléontologiques et biostratigraphiques. Ferrantia 62, Musée national d'histoire naturelle,
Luxembourg, 35-62.
20 = Kriwet J. 2008. A Late Jurassic carpetshark (Neoselachii, Orectolobiformes) from
southern Germany. Mesozoic Fishes 4 - Homology and Phylogeny, 443-454, 5 figs.
21 = Klug S, Kriwet J. 2013b. An offshore fish assemblage (Elasmobranchii, Actinopterygii)
from the Late Jurassic of NE Spain. Paläontologische Zeitschrift 87, 235-257.
22 = Hübner T, Müller A. 2010. Selachian teeth from Campanian sediments (Upper
Cretaceous) of the Münsterland Cretaceous Basin (NW-Germany). Paläontologische
Zeitschrift 84, 437-455.
23 = Bourdon J, Everhart MJ. 2010. Occurrence of the extinct Carpet shark,
Orectoloboides, in the Dakota Formation (Late Cretaceous; Middle Cenomanian) of
Kansas. Transactions of the Kansas Academy of Science 113, 237-242.
24 = Underwood CJ, Cumbaa SL. 2010. Chondrichthyans from a Cenomanian (Late
Cretaceous) bonebed, Saskatchewan, Canada. Palaeontology 53, 903-944.



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New images at shark-references:

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Please support shark-references and send your images to: info@shark-references.com

Missing papers:

Many thanks to all friends of shark-references, who send me some missing papers last month!

Shark-References would kindly like to ask you for your contribution to this project.

At the moment I search e.g. the following papers:

Journal of Ichthyology:
PINCHUK, V.I. & PERMITIN, Y.Y. 1970 New data on dogfish sharks of the Family Squalidae in the southeastern Atlantic. Journal of Ichthyology, 10 (3): 273-276
RASS, T.S. & LINDBERG, G.U. 1971 Modern concepts of the classification of living fishes. Journal of Ichthyology, 11: 302-319
GUBANOV, Y.P. 1972 On the biology of the thresher shark (Alopias vulpinus (Bonnaterre)) in the northwest Indian Ocean. Journal of Ichthyology, 12 (4): 591-600, figs 1-2, tabs 1-3
DOMANEVSKIY, L.N. 1975 The Frill Shark, Chlamydoselachus anguineus, from the Cape Blanc Area (Central Eastern Atlantic). Journal of Ichthyology, 15 (6): 1000-1002
GUBANOV, Y.P. & GRIGOREV, V.N. 1975 Distribution and biology of the blue shark Prionace glauca (Carcharhinidae) of the Indian Ocean. Journal of Ichthyology, 15: 37-43
GUBANOV, Y.P. 1976 The first catch of a tagged thresher shark (Alopias vulpinus). Journal of Ichthyology, 16 (3): 497-498
SVETLOV, M.F. 1978 The porbeagle, Lamna nasus, in Antarctic waters. Journal of Ichthyology, 18 (5): 850-851
GUBANOV, Y.P. 1978 The reproduction of some species of pelagic sharks from the equatorial zone of the Indian Ocean. Journal of Ichthyology, 18: 781-792
MYAGKOV, N.A. & KONDYURIN, V.V. 1978 Reproduction of the catshark Apristurus saldanha. Journal of Ichthyology, 4: 627-628
KONSTANTINOV, K.G. & NIZOVTSEV, G.P. 1979 The basking shark, Cetorhinus maximus, in Kandalaksha Bay of the White Sea. Journal of Ichthyology, 19 (1): 155-156
PIOTROVSKIY, A.S. & PRUT'KO, V.G. 1980 The occurrence of the goblin shark, Scapanorhynchus owstoni (Chondrichthyes, Scapanorhynchidae) in the Indian Ocean. Journal of Ichthyology, 20 (1): 124-125
TUMOKHIN, I.G. 1980 Discovery of the frill shark, Chlamydoselachus anguineus, in the southwest Indian Ocean.  Journal of Ichthyology, 20 (1): 125-126
PINCHUK, V.I. 1981 Mistaken identification of the so-called "dogfish", a member of the gray shark genus, Carcharhinus , from open waters of world oceans. Journal of Ichthyology, 21 (5): 115-117
LITVINOV, F.F. 1982 Two forms of teeth in Blue shark, Prionace glauca (Carcharhinidae). Journal of Ichthyology, 22 (4): 154-156
LITVINOV, F.F. & AGAPOV, S.N. & KATALIMOV, V.G. & MIRONOV, S.G. 1983 Rate of tooth Replacement in Blue Shark, Prionace glauca (Carcharhinidae), in relation to Feeding. Journal of Ichthyology, 23 (1): 143-145
KONDYURIN, V.V. & MYAGKOV, N.A. 1983 Catches of newborn Greenland shark, Somniosus microcephalus (Bloch and Schneider) (Dalatiidae). Journal of Ichthyology, 23 (6): 140-141
MYAGKOV, N.A. 1984 Unusual brain structure of luminous shark, Isistius brasiliensis (Dalatiidae). Journal of Ichthyology, 24 (2): 109-112
GUBANOV, E.P. 1985 Presence of the sharp tooth sand shark, Odontaspis ferox (Odontaspididae), in the open waters of the Indian Ocean. Journal of Ichthyology, 25 (2): 156-158
PARIN, N.V. & KOTLYAR, A.N. 1985 Electric rays of the genus Torpedo in open waters of the eastern south Pacific Ocean. Journal of Ichthyology, 26 (1): 1-12
GOLOVAN, G.A. & PAKHORUKOV, N.P. 1986 New records of rare species of cartilaginous fishes.  Journal of Ichthyology, 26: 117-120
MYAGKOV, N.A. & KONDYURIN, V.V. 1986 Dogfishes Squalus (Squalidae), of the Atlantic Ocean and comparative notes on the species of this genus from other regions. Journal of Ichthyology, 27 (1): 1-18
IVANOV, A. 1986 A new capture of the rare catshark, Apristurus longicephalus (Scyliorhinidae). Journal of Ichthyology, 27 (1): 147-149
IVANOV, A. 1987 On the distribution of the bigeye thresher shark, Alopias superciliosus, in the Pacific Ocean. Journal of Ichthyology, 26 (5): 121-122, fig
GUSHCHIN, A.V. & SUKHOVERSHIN, V.V. & KONOVALENKO, I.I. & SUKHORUKOVA, V.S. 1987 On the capture of the polar shark genus Somniosus (Squalidae) in the Southern Hemisphere. Journal of Ichthyology, 27 (1): 115-117
SHCHERBACHEV, Y.N. 1987 Preliminary list of thalassobathyal fishes of the tropical and subtropical waters of the Indian Ocean. Journal of Ichthyology, 27 (2): 37-46
MYAGKOV, N.A. 1987 External structure of the cephalic brain of the pelagic shark, Squaliolus laticaudus. Journal of Ichthyology, 27 (6): 125-127
GUBANOV, E.P. 1988 Morphological characteristics of the requiem shark, Carcharinus obscurus , of the Indian Ocean. Journal of Ichthyology, 28 (6): 68-73
KASHKIN, N.I. 1989 Mesopelagic ichthyofauna of the southwestern Pacific. Journal of Ichthyology, 29 (3): 116-127, tabs 1-4
BERESTOVSKIY, E.G. 1990 Feeding in the skates, Raja radiata and Raja fyllae, in the Barents and Norwegian seas. Journal of Ichthyology, 29 (8): 88-96
MANILO, L.G. 1993 New reports of fish on the shelf and upper slope of the Western Indian Ocean. Journal of Ichthyology, 33 (1): 128-136
BLAGODEROV, A.I. 1994 Seasonal distribution and some notes on the biology of salmon shark (Lamna ditropis) in the northwestern Pacific Ocean. Journal of Ichthyology, 34 (2): 115-121, figs 1-2, table
ROMANOV, E.V. & SAMOROV, V.V. 1994 On discoveries of the crocodile shark, Pseudocarcharias kamoharai (Pseudocarchariidae), in the Equatorial Indian Ocean. Journal of Ichthyology, 34 (4): 155-157
SAVELEV, S.V. & CHERNIKOV, V.P. 1994 The oceanic whitetip shark, Carcharhinus longimanus, and its use of aerial olfaction in search for food. Journal of Ichthyology, 34 (6): 38-47
PSHENICHNOV, L.K. 1997 A new record for subantarctic fish fauna species of shark Squalus acanthias (Squalidae). Journal of Ichthyology, 37 (8): 678-679
SOKOLOVSKAYA, T.G. & SOKOLOVSKII, A.S. & SOBOLEVSKII, E.I. 1998 A list of fishes of Peter the Great Bay (the Sea of Japan). Journal of Ichthyology, 38 (1): 1-11
DOLGANOV, V.N. 1999 Geographical and bathymetric distribution of the skates of the Rajidae family in the Far Eastern Seas of Russia and adjacent waters. Journal of Ichthyology, 39 (4): 340-342
NOVIKOV, N.P. 2002 Ecology of the ratfish Hydrolagus africanus (Gilchrist) from the Madagascar and Mozambique submarine ridges. Journal of Ichthyology, 42 (3): 271-274
MANILO, L.G. & BOGORODSKY, S.V. 2003 Taxonomic composition, diversity and distribution of coastal fishes of the Arabian Sea. Journal of Ichthyology, 43 (1): 75-149
KUKUEV, E.I. 2006 The second capture of a rare shark Scymnodalatias garricki (Dalatiidae) on a rise of the South Azores Complex and additional data on the distribution of S. albicauda. Journal of Ichthyology, 46 (9): 811-814
POLTEV, Y.N. & SHEIKO, B.A. 2007 Capture of Hydrolagus cf. purpurescens (Gilbert, 1905) (Chimaeriformes: Chimaeridae) off southeastern Sakhalin. Journal of Ichthyology, 47 (8): 611-619
PARIN, N.V. & KOTLYAR, A.N. 2007 On finding of shark of the genus Somniosus (Squalidae) at the submarine ridge of Nazca (southeastern Pacific). Journal of Ichthyology, 47 (8): 669-672
PARIN, N.V. & TIMOKHIN, I.G. & NOVIKOV, N.P. & SHCHERBACHEV, Y.N. 2008 On the composition of talassobathyal ichthyofauna and commercial productivity of Mozambique Seamount (the Indian Ocean). Journal of Ichthyology, 48 (5): 361-366
KUKUEV, E.I. & PAVLOV, V.P. 2008 The first case of mass catch of a rare frill shark Chlamydoselachus anguineus over a seamount of the Mid-Atlantic Ridge. Journal of Ichthyology, 48 (8): 676-678
PROKOFIEV, A.M. & KUKUEV, E.I. 2009 New findings of rare fish species from families Mitsukurinidae (Chondrichthyes), Muraenidae, Lophiidae, Macrouridae, and Psychrolutidae (Teleostei) on raises of the atlantic ocean with the description of Gymnothorax walvisensis sp. nova. Journal of Ichthyology, 49 (3): 215-227
Annals and Magazine of Natural History
COUCH, J. 1838 Description of a species of ray-fish, not hitherto included in the British fauna. Annals and Magazine of Natural History, (Series 1), 2: 71-73
CHARLESWORTH, E. 1839 On the remains of a species of Hybodus from Lyme Regis. Annals and Magazine of Natural History, (Series 3): 242-248
Aqua, International Journal of Ichyology
LASSO, C.A. & RIAL, B.A. & LASSO-ALCALA, O. 1997 Notes on the biology of the freshwater stingrays Paratrygon aiereba (Müller & Henle, 1841) and Potamotrygon orbignyi (Castelnau, 1855) (Chondrichthyes: Potamotrygonidae) in the Venezuelan Llanos. Aqua, International Journal of Ichyology, 2 (3): 39-50
HUMAN, B.A. 2011 Description of a unique catshark egg capsule (Chondrichthyes: Scyliorhinidae) from the North West Shelf, Western Australia. Aqua, International Journal of Ichyology, 17 (4): 199-209
Canadian Journal of Earth Sciences
MASSON, A.G. & RUST, B.R. 1984 Freshwater shark teeth as paleoenvironmental indicators in the Upper Pennsylvanian Morien Group of the Sydney Basin, Nova Scotia Canadian Journal of Earth Sciences, 21 (10): 1151-1155
FIELITZ, C. 1996 A Late Cretaceous (Turonian) ichthyofauna from Lac des Bois, Northwest Territories, Canada, with paleobiogeographic comparisons with Turonian ichthyofaunas of the Western Interior Seaway.Canadian. Canadian Journal of Earth Sciences, 33: 1375-1389
Transactions of the Royal Society of Edinburgh
TRAQUAIR, R.H. 1903 On the distribution of fossil fish-remains in the Carboniferous rocks of the Edinburgh district. Transactions of the Royal Society of Edinburgh, 40 (3): 687-707
WESTOLL, T.S. 1947 The Paired Fins of Placoderms. Transactions of the Royal Society of Edinburgh, 61 (2): 381-398
DICK, J.R.F. 1978 On the Carboniferous shark Tristychius arcuatus Agassiz from Scotland. Transactions of the Royal Society of Edinburgh, 70 (4): 63-109
PATON, R.L. 1993 Elasmobranch fishes from the Viséan of East Kirkton, West Lothian, Scotland. Transactions of the Royal Society of Edinburgh, 84 (3-4): 329-330
COATES, M.I. & SEQUEIRA, S.E.K. 1998 The braincase of a primitive shark. Transactions of the Royal Society of Edinburgh, Earth Sciences, 89: 63-85


Please support www.shark-references.com and send missing papers (not listed papers or papers without the infosymbol) to juergen.pollerspoeck@shark-references.com.






EEA 2014 European Elasmobranch Association – Annual Scientific Conference 18 years on – Prepared for the Future

The Dutch Elasmobranch Society is proud to host the 18th Annual Scientific Conference of the European Elasmobranch Association from 7th to 9th November 2014 at the Van Hall Larenstein University of Applied Sciences in Leeuwarden, The Netherlands.

Elasmobranchs are increasingly included in high level policy agreements. The conference will provide a platform for those involved in international science and policy and aims to help coordinate the information necessary for the development and implementation of management measures for rays and sharks in European waters. EEA 2014 will be of interest to all those who are involved in the study, management and conservation of chondrichthyans (sharks, skates, rays and chimaeras).

The three-day conference will include theme sessions on policy making, restoration measures, integrated management, husbandry and captive management, tagging and other subjects. The collection, availability and sharing of data will be an underlying theme. There will be plenary talks, a poster session and opportunities for networking and socialising. In addition an excursion will be offered on the last day.

Leeuwarden is a vibrant town with a charming centre and a wide selection of hotels and restaurants. It is the capital city of the northern Province of Fryslân and will be the Cultural Capital of Europe in 2018. There is a twice hourly train service from Schiphol (one direct, one with one connection) and it is easy to reach by road.

See www.elasmobranch.nl/EEA2014 for preliminary details on the conference.

Contact: eea2014@elasmobranch.nl

New described species/Taxonomic News:


Rhinobatos whitei LAST, CORRIGAN & NAYLOR, 2014, adolescent male holotype, 597 mm TL (PNM 15189). Dorsal view (fresh from copy of printed image); © Peter Last, CSIRO - Australian National Fish Collection
LAST, P.R. & CORRIGAN, S. & NAYLOR, G.J.P. (2014):  Rhinobatos whitei, a new shovelnose ray (Batoidea: Rhinobatidae) from the Philippine Archipelago. Zootaxa, 3872 (1): 31-47
New species: Rhinobatos whitei
Abstract: A new shovelnose ray, Rhinobatos whitei sp. nov., is described from material collected at fish markets of the southern Philippines. This ray was first formally indentified as an undescribed species more than a decade ago as part of a WWF funded survey of sharks and rays of the Philippines. It was considered to be most closely related to another shovelnose ray found nearby in the western North Pacific, R. schlegelii, but differs from that species in body shape and aspects of coloration, meristics and morphometry. It differs from all other shovelnose rays of the region in its NADH2 sequence, clustering together with an Indonesian species R. jimbaranensis, and another undescribed species from Borneo.

WHITE, W.T. & VAZ, D.F.B. & HO, H.-C. & EBERT, D.A. & DE CARVALHO, M.R. & CORRIGAN, S. & ROCHEL, E. & DE CARVALHO, M. & TANAKA, S. & NAYLOR, G.J.P. (2014):  Redescription of Scymnodon ichiharai Yano and Tanaka 1984 (Squaliformes: Somniosidae) from the western North Pacific, with comments on the definition of somniosid genera. Ichthyological Research, in press
Redescribed species: Scymnodon ichiharai
Abstract: The somniosid species Scymnodon ichiharai Yano and Tanaka 1984 is redescribed based on the type specimens and additional material from Taiwan and Japan. The range of this species is extended to include Taiwanese waters. Although recently allocated to the genus Zameus, this species is very similar to Scymnodon plunketi from the southern Indo-West Pacific; our molecular evidence indicates that these species are sister-taxa. The genus Proscymnodon Fowler 1934 is placed in the synonymy of Scymnodon Barboza du Bocage and de Brito Capello 1864 and new generic definitions are provided for Scymnodon and the closely related Zameus. The genusScymnodon is considered to include at least the species S. ichiharaiS. plunketi, and S. ringens;Proscymnodon macracanthus (Regan 1906) is allocated to Scymnodon, but its validity is uncertain. The genus Zameus is considered to be monotypic, containing only the species Z. squamulosus.


Parasquatina zitteli nov. sp. (Elasmobranchii: Orectolobiformes) aus dem Maastricht von Oberbayern (Gerhartsreiter Schichten, Siegsdorf) und Bemerkungen zur Verbreitung der Ordnung Orectolobiformes. [Parasquatina zitteli nov. sp. (Elasmobranchii: Orectolobiformes) from the Maastrichian of Upper Bavaria (Gerhartsreiter Ditch, Siegsdorf), with remarks on the distribution of the order Orectolobiformes.] Zitteliana, A54: 147-164
New species: Parasquatina zitteli
Abstract: The genus Parasquatina is documented for the first time from the Gerhartsreiter Ditch near Siegsdorf (Upper Bavaria, Germany). The fossils are interpreted as a new species, for which the name Parasquatina zitteli nov. sp. is proposed, because they differ in morphology from all other species in that genus, especially with regard to the labial apron, which is absent in P. zitteli. Assignment of the genus Parasquatina to the order Squatiniformes is reversed, and the genus now placed in the order Orectolobiformes. In addition, a survey of recent records of Parasquatina is given, together with an up-to-date overview of the stratigraphic distribution of the order Orectolobiformes, and resulting questions are discussed.


PLEASE send your new papers tojuergen.pollerspoeck@shark-references.com!

New Paper


Recent Papers:

AKHILESH K.V. & BINEESH, K.K. & GOPALAKRISHNAN, A. & JENA, J.K. & BASHEER, V.S. & PILLAI, N.G.K. (2014):  Checklist of Chondrichthyans in Indian waters. Journal of the Marine Biological Association of India, 56 (1): 109-120 http://dx.doi.org/10.6024/jmbai.2014.56.1.01750s-17
ÁNGEL, F. & QUINTEIRO, J. & REY-MÉNDEZ, M. (2014):  First record for the Caribbean Sea of the shark Echinorhinus brucus captured in Venezuelan waters.Marine Biodiversity Records, 7: e91  http://dx.doi.org/10.1017/S1755267214000967
BANDYOPADHYAY, P.R. & HELLUM, A.M. (2014):  Modeling how shark and dolphin skin patterns control transitional wall-turbulence vorticity patterns using spatiotemporal phase reset mechanisms. Scientific Reports, 4: 6650 http://dx.doi.org/10.1038/srep06650
BETHEA, D.M. & AJEMIAN, M.J. & CARLSON, J.K. & HOFFMAYER, E.R. & IMHOFF, J.L. & GRUBBS, R.D. & PETERSON, C.T. & BURGESS, G.H. (2014): Distribution and community structure of coastal sharks in the northeastern Gulf of Mexico. Environmental Biology of Fishes, in press  http://dx.doi.org/10.1007/s10641-014-0355-3
BIZZARRO, J.J. & BROMS, K.M. & LOGSDON, M.G. & EBERT, D.A. & YOKLAVICH, M.M. & KUHNZ, L.A. & SUMMERS, A.P. (2014):  Spatial Segregation in Eastern North Pacific Skate Assemblages. PLoS ONE, 9 (10): e109907 http://dx.doi.org/10.1371/journal.pone.0109907
CARDEÑOSA, D. & HYDE, J. & CABALLERO, S. (2014):  Genetic Diversity and Population Structure of the Pelagic Thresher Shark (Alopias pelagicus) in the Pacific Ocean: Evidence for Two Evolutionarily Significant Units. PLoS ONE, 9 (10): e110193 http://dx.doi.org/10.1371/journal.pone.0110193
CASTILLO-PÁEZ, A. & DEL RÍO-PORTILLA, M.A. & OÑATE-GONZÁLEZ, E. & ROCHA-OLIVARES, A. (2014):  The mitochondrial genome of the banded guitarfish, Zapteryx exasperata (Jordan and Gilbert, 1880), possesses a non-coding duplication remnant region. Mitochondrial DNA, in press http://dx.doi.org/10.3109/19401736.2014.958721
CASTILLO-PÁEZ, A. & DEL RÍO-PORTILLA, M.A. & ROCHA-OLIVARES, A. (2014):  The complete mitochondrial genome of the giant electric ray, Narcine entemedor (Elasmobranchii: Torpediniformes). Mitochondrial DNA, in press http://dx.doi.org/10.3109/19401736.2014.963800
CHEN, X. & XIANG, D. & PENG, X. & AI, W. & CHEN, H. (2014):  The complete mitochondrial genome of the spinner shark Carcharhinus brevipinna. Mitochondrial DNA, in press  http://dx.doi.org/10.3109/19401736.2014.958720
COLONELLO, J.H. & CORTÉS, F. & MASSA, A.M. (2014):  Species richness and reproductive modes of chondrichthyans in relation to temperature and fishing effort in the Southwestern Atlantic Shelf (34–54° S). Fisheries Research, 160: 8-17 http://dx.doi.org/10.1016/j.fishres.2014.04.015
DALY, R. & SMALE, M.J. & COWLEY, P.D. & FRONEMAN, P.W. (2014): Residency Patterns and Migration Dynamics of Adult Bull Sharks (Carcharhinus leucas) on the East Coast of Southern Africa. PLoS ONE, 9 (10): e109357 http://dx.doi.org/10.1371/journal.pone.0109357
DAVIDSON, B.C. & NEL, W. & RAIS, A. & NAMDARIZANDI, V. & VIZARRA, S. & CLIFF, G. (2014):  Comparison of total lipids and fatty acids from liver, heart and abdominal muscle of scalloped (Sphyrna lewini) and smooth (Sphyrna zygaena) hammerhead sharks. Springerplus, 3: 521  http://dx.doi.org/10.1186/2193-1801-3-521
DRIGGERS, W.B. & DAVIS, K.S. & MOORE, C. & CARLSON, J.K. (2014):  New record of a goblin shark Mitsukurina owstoni (Lamniformes: Mitsukurinidae) in the western North Atlantic Ocean. Marine Biodiversity Records, 7: e96 http://dx.doi.org/10.1017/S1755267214000876
DULVY N.K. & DAVIDSON L.N.K. & KYNE P.M. & SIMPFENDORFER C.A. & HARRISON L.R. & CARLSON J.K. & FORDHAM S.V. (2014):  Ghosts of the coast: global extinction risk and conservation of sawfishes. Aquatic Conservation: Marine and Freshwater Ecosystems, in press  http://dx.doi.org/10.1002/aqc.2525
EGEBERG, C.A. & KEMPSTER, R.M. & THEISS, S.M. & HART, N.S. & COLLIN, S.P. (2014):  The distribution and abundance of electrosensory pores in two benthic sharks: a comparison of the wobbegong shark, Orectolobus maculatus, and the angel shark, Squatina australis. Marine and Freshwater Research, 65 (11): 1003–1008 http://dx.doi.org/10.1071/MF13213
FUSS, T. & SCHLUESSEL, V. (2014):  Something worth remembering: visual discrimination in sharks. Animal Cognition, in press  http://dx.doi.org/10.1007/s10071-014-0815-3
GODARD, B.G. & COOLEN, M. & LE PANSE, S. & GOMBAULT, A. & FERREIRO-GALVE, S. & LAGUERRE, L. & LAGADEC, R. & WINCKER, P. & POULAIN, J. & DA SILVA, C. & KURAKU, S. & CARRE, W. & BOUTET, A. & MAZAN, S. (2014): Mechanisms of endoderm formation in a cartilaginous fish reveal ancestral and homoplastic traits in jawed vertebrates. Biology Open: BIO20148037 http://dx.doi.org/10.1242/bio.20148037
HEINRICH, D.D.U. &, RUMMER, J.L. &, MORASH, A.J. &, WATSON, S.-A. &, SIMPFENDORFER, C.A. &, HEUPEL, M.R. &, MUNDAY, P.L. (2014):  A product of its environment: the epaulette shark (Hemiscyllium ocellatum) exhibits physiological tolerance to elevated environmental CO2. Conservation Physiology, 2 (1): cou047 http://dx.doi.org/10.1093/conphys/cou047
HORNING, M. & MELLISH, J.-A.E. (2014):  In cold blood: evidence of Pacific sleeper shark (Somniosus pacificus) predation on Steller sea lions (Eumetopias jubatus) in the Gulf of Alaska. Fishery Bulletin, 112 (4): 297-310  http://dx.doi.org/10.7755/FB.112.4.6
HOSSAIN, M.A. & THOMPSON, B.S. & CHOWDHURY, G.W. & MOHSANIN, S. & FAHAD, Z.H. & KOLDEWEY, H.J. & ISLAM, M.A. (2014):  Sawfish exploitation and status in Bangladesh. Aquatic Conservation: Marine and Freshwater Ecosystems, in press  http://dx.doi.org/10.1002/aqc.2466
HYODO, S. & KAKUMURA, K. & TAKAGI, W. & HASEGAWA, K. & YAMAGUCHI, Y. (2014):  Morphological and functional characteristics of the kidney of cartilaginous fishes: with special reference to urea reabsorption. American Journal of Physiology - Regulatory, Integrative and Comparative Physiology, in press http://dx.doi.org/10.1152/ajpregu.00033.2014
JABADO, R.W. & AL GHAIS, S.M. & HAMZA, W. & SHIVJI, M.S. & HENDERSON, A.C. (2014):  Shark diversity in the Arabian/Persian Gulf higher than previously thought: insights based on species composition of shark landings in the United Arab Emirates. Marine Biodiversity, in press  http://dx.doi.org/10.1007/s12526-014-0275-7
KABASAKAL, H. (2014):  The status of the great white shark (Carcharodon carcharias) in Turkey’s waters. Marine Biodiversity Records, 7: e109 http://dx.doi.org/10.1017/S1755267214000980
KHADEEJA, A. & SINAN, H. (2014):  Shark ban in its infancy: Successes, challenges and lessons learned. Journal of the Marine Biological Association of India, 56 (1): 34-40  http://dx.doi.org/10.6024/jmbai.2014.56.1.01750s-05
LAST, P.R. & CORRIGAN, S. & NAYLOR, G.J.P. (2014):  Rhinobatos whitei, a new shovelnose ray (Batoidea: Rhinobatidae) from the Philippine Archipelago. Zootaxa, 3872 (1): 31-47  http://dx.doi.org/10.11646/zootaxa.3872.1.3
LEVINE, M. & COLLIER, R.S. & RITTER, E. & FOUDA, M. & CANABAL, V. (2014): Shark Cognition and a Human Mediated Driver of a Spate of Shark Attacks. Open Journal of Animal Sciences, 4 (5): 50200  http://dx.doi.org/10.4236/ojas.2014.45033
LI, W. & DAI, X. & XU, Q. & WU, F. & GAO, C. & ZHANG, Y. (2014):  The complete mitochondrial genome sequence of Oceanic whitetip shark, Carcharhinus longimanus (Carcharhiniformes: Carcharhinidae). Mitochondrial DNA, in press http://dx.doi.org/10.3109/19401736.2014.963807
MCCLUSKY, L.M. & SULIKOWSKI, J. (2014):  The epigonal organ and mature pole of the testis in the recreationally fished blue shark (Prionace glauca): histochemico-functional correlates. Journal of Anatomy, in press  http://dx.doi.org/10.1111/joa.12242
MONTEALEGRE-QUIJANO, S. & CARDOSO, A.T.C. & SILVA, R.Z. & KINAS, P.G. & VOOREN, C.M. (2014):  Sexual development, size at maturity, size at maternity and fecundity of the blue shark Prionace glauca (Linnaeus, 1758) in the Southwest Atlantic.Fisheries Research, 160: 18–32  http://dx.doi.org/10.1016/j.fishres.2014.03.003
MUNROE, S.E.M. & SIMPFENDORFER, C.A. & HEUPEL, M.R. (2014):  Habitat and space use of an abundant nearshore shark, Rhizoprionodon taylori. Marine and Freshwater Research, 65 (11): 959–968  http://dx.doi.org/10.1071/MF13272
OLIN, J.A. & POULAKIS, G.R. & STEVENS, P.W. & DEANGELO, J.A. & FISK, A.T. (2014):  Preservation Effects on Stable Isotope Values of Archived Elasmobranch Fin Tissue: Comparisons between Frozen and Ethanol-Stored Samples. Transactions of the American Fisheries Society, 143 (6): 1569-1576 http://dx.doi.org/10.1080/00028487.2014.954055
ORLOV, A.M. & BAITALYUK, A.A. (2014):  Spatial Distribution and Features of Biology of Pacific Sleeper Shark Somniosus pacificus in the North Pacific. Journal of Ichthyology, 54 (8): 526-546  http://dx.doi.org/10.1134/S0032945214040080
PORTNOY, D.S. & HOLLENBECK, C.M. & BELCHER, C.N. & DRIGGERS, W.B. & FRAZIER, B.S. & GELSLEICHTER, J. & GRUBBS, R.D. & GOLD, J.R. (2014): Contemporary population structure and post-glacial genetic demography in a migratory marine species, the blacknose shark, Carcharhinus acronotus. Molecular Ecology, in press  http://dx.doi.org/10.1111/mec.12954
RIGBY, C.L. & WEDDING, B.B. & GRAUF, S. & SIMPFENDORFER, C.A. (2014): The utility of near infrared spectroscopy for age estimation of deepwater sharks. DeepSea Research Part I: Oceanographic Research Papers, in press http://dx.doi.org/10.1016/j.dsr.2014.09.004
RODRÍGUEZ-CABELLO, C. & SÁNCHEZ, F. (2014):  Is Centrophorus squamosus a highly migratory deep-water shark? Deep Sea Research Part I: Oceanographic Research Papers, 92: 1-10  http://dx.doi.org/10.1016/j.dsr.2014.06.005
SCHLUESSEL, V. & GILES, J. & KYNE, P.M. (2014):  Notes on female reproductive biology and embryos of the brown guitarfish Rhinobatos schlegelii from the Penghu Islands, Taiwan. Ichthyological Research, in press  http://dx.doi.org/10.1007/s10228-014-0431-x
SEN, S. & DASH, G. & BHARADIYA, S.A. (2014):  First record of blue-spotted stingray, Neotrygon kuhlii from Gujarat, north-west coast of India. Marine Biodiversity Records, 7: e81  http://dx.doi.org/10.1017/S1755267214000852
STEHFEST, K.M. & PATTERSON, T.A. & BARNETT, A. & SEMMENS, J.M. (2014): Markov models and network analysis reveal sex-specifi c differences in the space-use of a coastal apex predator. Oikos, in press  http://dx.doi.org/10.1111/oik.01429
TAO, Z. & BULLARD, S.A. & ARIAS, C.R. (2014):  Diversity of bacteria cultured from the blood of lesser electric rays caught in the northern gulf of Mexico. Journal of Aquatic Animal Health, 26 (4): 225-232 http://dx.doi.org/10.1080/08997659.2014.922513
TOMITA, T. & NAKAMURA, M. & SATO, K. & TAKAOKA, H. & TODA, M. & KAWAUCHI, J. & NAKAYA, K. (2014):  Onset of Buccal Pumping in Catshark Embryos: How Breathing Develops in the Egg Capsule. PLoS ONE, 9 (10): e109504 
TRINNIE, F.I. & WALKER, T.I. & JONES, P.L. & LAURENSON, L.J. (2014): Regional differences in the reproductive parameters of the sparsely-spotted stingaree, Urolophus paucimaculatus, from south-eastern Australia. Marine and Freshwater Research, 65 (11): 943-958  http://dx.doi.org/10.1071/MF13275
VERAS, D.P. & HAZIN, F.H.V. & BRANCO, I.S.L. & TOLOTTI, M.T. & BURGESS, G.H. (2014):  Reproductive biology of the pelagic stingray, Pteroplatytrygon violacea (Bonaparte, 1832), in the equatorial and south-western Atlantic Ocean. Marine and Freshwater Research, 65 (11): 1035–1044  http://dx.doi.org/10.1071/MF13008
WCISEL, M. & O’RIAIN, M.J. & DE VOS, A. & CHIVELL, W. (2014):  The role of refugia in reducing predation risk for Cape fur seals by white sharks. Behavioral Ecology and Sociobiology, in press  http://dx.doi.org/10.1007/s00265-014-1825-5
WHITE, W.T. & VAZ, D.F.B. & HO, H.-C. & EBERT, D.A. & DE CARVALHO, M.R. & CORRIGAN, S. & ROCHEL, E. & DE CARVALHO, M. & TANAKA, S. & NAYLOR, G.J.P. (2014):  Redescription of Scymnodon ichiharai Yano and Tanaka 1984 (Squaliformes: Somniosidae) from the western North Pacific, with comments on the definition of somniosid genera. Ichthyological Research, in press http://dx.doi.org/10.1007/s10228-014-0430-y
WYFFELS, J. & KING, B.L. & VINCENT, J. & CHEN, C. & WU, C.H. & POLSON, S.W. (2014):  SkateBase, an elasmobranch genome project and collection of molecular resources for chondrichthyan fishes. F1000Research, 3: 191 http://dx.doi.org/10.12688/f1000research.4996.1
ZHANG, X. & ZHAO, H. & CHEN, Y. & LUO, H. & YAO, B. (2014):  Molecular characterization of a CpTRIM35-like protein and its splice variants from whitespotted bamboo shark (Chiloscyllium plagiosum). Biochemical and Biophysical Research Communications, in press  http://dx.doi.org/10.1016/j.bbrc.2014.09.139




BULGUROĞLU, S.Y. & KORUN, J. & GÖKOĞLU, M. & ÖZVAROL, Y. (2014):  The marine leech Stibarobdella moorei (Oka, 1910) (Hirudinea, Piscicolidae) parasitic on the thornback ray Raja clavata Linnaeus, 1758 and angelshark Squatina squatina (Linnaeus, 1758) in Antalya Bay, Mediterranean Sea of Turkey. Helminthologia, 51 (3): 250-252  http://dx.doi.org/10.2478/s11687-014-0237-4 



EHRET, D.J. & EBERSOLE, J.A. (2014):  Occurrence of the megatoothed sharks (Lamniformes: Otodontidae) in Alabama, USA. PeerJ, 2: e625 http://dx.doi.org/10.7717/peerj.625
EHRET, D.J. & EBERSOLE, J.A. (2014):  Occurrence of the Megatoothed sharks (Lamniformes:Otodontidae) in Alabama, USA. Abstract. PeerJ PrePrints, 2: e517v1 
FISCHER, J. (2014):  Palaeoecology, migration behavior, and reproductive pattern of Palaeozoic to Mesozoic freshwater sharks revealed by stable and radiogenic isotopes. Abstract. Geologica Saxonica, 60 (2): 299-300
LADWIG, J. (2014):  Haizähne der Art Squalicorax kaupi (AGASSIZ, 1843) (Lamniformes, Anacoracidae) aus Ignaberga (Schonen, Schweden). Arbeitskreis Paläontologie Hannover, 42: 101–103
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Sharks that hide in coral reefs may be safe from acidifying oceans

October 15, 2014
Oxford University Press (OUP)
The epaulette shark displays physiological tolerance to elevated carbon dioxide in its environment after being exposed to carbon dioxide levels equivalent to those that are predicted for their natural habitats in the near future.

A study published online today in the journal Conservation Physiologyhas shown that the epaulette shark (Hemiscyllium ocellatum) displays physiological tolerance to elevated carbon dioxide (CO₂) in its environment after being exposed to CO₂ levels equivalent to those that are predicted for their natural habitats in the near future.

read the full story:


News from ASIS Australian Shark Information System:

Looking for specific data about shark and ray species and fisheries? 
ASIS will be available for download as ASIS Desktop, a Microsoft Access database file you use on your own computer as a 'desktop' version. Use ASIS Desktop to find and view ASIS data about specific sharks and rays, and/or fisheries. 
Help make sure ASIS data is up to date with your research and information!
 Download Data Release 1 sharks & fisheries
 Download Data Release 2 sharks species data
Data Release 1 is an Excel spreadsheet summarising 137 Australian fisheries and their interactions with sharks and rays. Fishery managers and researchers are invited to download the spreadsheet and validate the data. Please note:
Some records are blank for the management field. This is usually where there are multiple sectors of the fishery and the management for all sectors is described in the record for All sectors. These are highlighted in green.
The data in the management field is intentionally limited to details about arrangements that are most likely to affect sharks and rays. 
Data Release 2 - shark species data is an Excel spreadsheet of species data for 28 species from the Chlamydoselachidae to Centrophoridae. Fishery managers and researchers are invited to download the spreadsheet and validate the data, and to nominate expert contacts (self-nominations are accepted) for each species.

For more information please visit: http://www.sharkreportcard.org/#!asis-data/ca4p


When Did Carcharocles megalodon Become Extinct?

image: Tooth of megalodon (Carcharocles megalodon) with ruler for scale

- Carcharocles megalodon (“Megalodon”) is the largest shark that ever lived.
- The results of this study suggest that C. megalodon went extinct around 2.6 Ma.

open access:
Pimiento C, Clements CF (2014) When Did Carcharocles megalodon Become Extinct? A New Analysis of the Fossil Record. PLoS ONE 9(10): e111086