Helodus sp.
Classification: Holocephali Helodontiformes Helodontidae
	Types
Helodus sp.
	
	
	
Helodus sp.
Description:
Citation: Helodus sp. : In: Database of fossil elasmobranch teeth www.shark-references.com, World Wide Web electronic publication, Version 11/2025
		Remarks
shark-references Species-ID=8139;
		
		shark-references Species-ID=8139;
		References
		
	
		
		
			
Rise and diversification of chondrichthyans in the Paleozoic. Paleobiology, 50(2), 271–284
DOI: 10.1017/pab.2024.1
	
	
		
		
			
Sharks in the dark: Paleontological resource inventory reveals multiple successive Mississippian Subperiod cartilaginous fish (Chondrichthyes) assemblages within Mammoth Cave National Park, Kentucky. Parks Stewardship Forum, 40(1), 53–67
DOI: 10.5070/P540162921
	
	
		
		
			
A late Permian ichthyofauna from the Zechstein Basin, Lithuanian–Latvian Region. Journal of Iberian Geology, 47, 461–481
DOI: 10.1007/s41513-020-00135-y
	
	
		
		
			
Pennsylvanian brachiopod, fish and conodont faunas from the Caliza Masiva (San Emiliano Formation) at the Mina Profunda area, Cantabrian Zone, NW Spain. Geobios, 59, 91–106
DOI: 10.1016/j.geobios.2020.03.004
	
	
		
		
			
The Holocephalans (Chondrichthyes) of the Mississippian (Visean) Redwall Limestone, Grand Canyon National Park, Arizona. New Mexico Museum of Natural History and Science, Bulletin, 82, 141–144
	
	
		
		
			
Late Pennsylvanian fish assemblage from the Robledo Mountains and new records of Paleozoic chondrichthyans in New Mexico, USA. Bulletin of Geosciences, 94(2), 235–255
DOI: 10.3140/bull.geosci.1741
	
	
		
		
			
The first microfossil record of ichthyofauna from the Naujoji Akmenė Formation (Lopingian), Karpėnai Quarry, North Lithuania. Geological Quarterly, 61(3), 602–610
	
	
		
		
			
Paleozoic fishes of New Mexico: a review. New Mexico Museum of Natural History and Science, Bulletin, 68, 51–64
	
	
		
		
			
Lower Permian Freshwater Sharks and Fishes of Texas and Oklahoma. published by the author himself, 118pp
	
	
		
		
			
Vertebrate biostratigraphy and biochronology of the upper Paleozoic Dunkard Group, Pennsylvania-West Virginia-Ohio, USA. International Journal of Coal Geology, 119, 79–87
DOI: 10.1016/j.coal.2013.04.007
	
	
		
		
			
Xenacanth Sharks and other Vertebrates from the Geraldine Bonebed, Lower Permian of Texas. New Mexico Museum of Natural History and Science, Bulletin, 60, 161–167
	
	
		
		
			
Sharks from the geraldine bonebed, lower permian of Texas [Abstract]. Journal of Vertebrate Paleontology, 32(Suppl. 1), 118
DOI: 10.1080/02724634.2012.10635175
	
	
		
		
			
A carboniferous chondrichthyan assemblage from residues within a Triassic karst system at Cromhall quarry, Gloucestershire, England. Palaeontology, 55(6), 1245–1263
DOI: 10.1111/j.1475-4983.2012.01193.x
	
	
		
		
			
Fish fossils from the Paleozoic Sly Gap Formation of southern New Mexico, USA. New Mexico Museum of Natural History and Science, Bulletin, 53, 52–70
	
	
		
		
			
Fishes and tetrapods in the Upper Pennsylvanian (Kasimovian) Cohn Coal Member of the Mattoon Formation of Illinois, United States: Systematics, Paleoecology, and Paleoenvironments. Palaios, 26(10), 639–657
DOI: 10.2110/palo.2010.p10-121r
	
	
		
		
			
Middle Palaeozoic microvertebrate assemblages and biogeography of East Gondwana (Australasia, Antarctica). Palaeoworld, 19(1–2), 37–54
DOI: 10.1016/j.palwor.2009.11.001
	
	
		
		
			
New specimens of xenacanth and hybodont sharks (Elasmobranchii: Xenacanthida and Hybodontoidea) from the Lower Permian of southwestern Oklahoma. Oklahoma Geology Notes, 63(4), 136–147
	
	
		
		
			
Fish Assemblages in the Tournaisian-Viséan environments of the East European Platform. In Strogen, P., Somerville, I. D. and Jones, G. L. (Eds.), Recent Advances in Lower Carboniferous Geology. Geol. Soc. Spec. Publ., 107, p. 387–415.
	
	
		
		
			
Vertebrate Microfossils from the Lueders Formation, Albany Group, and the faunal transition from the Wichita Group into the Clear Fork Group, Lower Permian of Texas. Modern Geology, 20, 371–382
	
	
		
		
			
Reworked Courceyan (Early Carboniferous) chondrichthyan remains from British Triassic cave deposits. In Turner, S.(Ed.) IGCP328: SDS The Gross Symposium, Göttingen 31 July – 7 Aug. 1993. Abstracts. 2pp.
	
	
		
		
			
Early Carboniferous Shark Remains from the Rockhampton District, Queensland. Memoirs of the Queensland Museum, 28(1) , 65–73
	
	
		
		
			
Clear Fork vertebrates and environments from the lower Permian of North-Central Texas. The Texas Journal of Science, 39(3): 253–266
	
	
		
		
			
Middle Palaeozoic elasmobranch remains from Australia. Journal of Vertebrate Paleontology, 2(2), 117–131
DOI: 10.1080/02724634.1982.10011923
	
	
		
		
			
Fossil fishes from the marine Late Palaeozoic of Holm Land-Amdrup Land, North-East Greenland. Meddelelser om Grønland, 195(9), 1–38
	
	
		
		
			
CHAPTER 26 Pisces In The Fossil Record. HARLAND, W.B. et al. (Eds.) 1967, London (Geological Society), pp: 627-683
	
	
		
		
			
Catalogue of Canadian fossil fishes. Life Sciences Contribution of the Royal Ontario Museum, 68, 1–154
	
	
		
		
			
Bradyodont sharks in the Permian of Western Australia. American Journal of Science, 243, 543–552
	
	
		
		
			
Fish remains from the Salem Limestone of Indiana. In Cumings E. R., Branson E. B., Beede, J. W. and Smith, E. A. (Eds.), The fauna of the Salem Limestone of Indiana. Rep. Indiana Dept. Geol. Natur. Resources, 30: 1376–1394
	
	
		
		
			
On some new species of fossil fish from the Millstone Grit rocks, with an amended list of genera and species. Proceedings of the Yorkshire Geological and Polytechnic Society, 15, 70-78
DOI: 10.1144/pygs.15.1.70
	
	
Rise and diversification of chondrichthyans in the Paleozoic. Paleobiology, 50(2), 271–284
DOI: 10.1017/pab.2024.1
Sharks in the dark: Paleontological resource inventory reveals multiple successive Mississippian Subperiod cartilaginous fish (Chondrichthyes) assemblages within Mammoth Cave National Park, Kentucky. Parks Stewardship Forum, 40(1), 53–67
DOI: 10.5070/P540162921

A late Permian ichthyofauna from the Zechstein Basin, Lithuanian–Latvian Region. Journal of Iberian Geology, 47, 461–481
DOI: 10.1007/s41513-020-00135-y

Pennsylvanian brachiopod, fish and conodont faunas from the Caliza Masiva (San Emiliano Formation) at the Mina Profunda area, Cantabrian Zone, NW Spain. Geobios, 59, 91–106
DOI: 10.1016/j.geobios.2020.03.004

The Holocephalans (Chondrichthyes) of the Mississippian (Visean) Redwall Limestone, Grand Canyon National Park, Arizona. New Mexico Museum of Natural History and Science, Bulletin, 82, 141–144

Late Pennsylvanian fish assemblage from the Robledo Mountains and new records of Paleozoic chondrichthyans in New Mexico, USA. Bulletin of Geosciences, 94(2), 235–255
DOI: 10.3140/bull.geosci.1741
The first microfossil record of ichthyofauna from the Naujoji Akmenė Formation (Lopingian), Karpėnai Quarry, North Lithuania. Geological Quarterly, 61(3), 602–610

Paleozoic fishes of New Mexico: a review. New Mexico Museum of Natural History and Science, Bulletin, 68, 51–64

Lower Permian Freshwater Sharks and Fishes of Texas and Oklahoma. published by the author himself, 118pp

Vertebrate biostratigraphy and biochronology of the upper Paleozoic Dunkard Group, Pennsylvania-West Virginia-Ohio, USA. International Journal of Coal Geology, 119, 79–87
DOI: 10.1016/j.coal.2013.04.007

Xenacanth Sharks and other Vertebrates from the Geraldine Bonebed, Lower Permian of Texas. New Mexico Museum of Natural History and Science, Bulletin, 60, 161–167

Sharks from the geraldine bonebed, lower permian of Texas [Abstract]. Journal of Vertebrate Paleontology, 32(Suppl. 1), 118
DOI: 10.1080/02724634.2012.10635175

A carboniferous chondrichthyan assemblage from residues within a Triassic karst system at Cromhall quarry, Gloucestershire, England. Palaeontology, 55(6), 1245–1263
DOI: 10.1111/j.1475-4983.2012.01193.x

Fish fossils from the Paleozoic Sly Gap Formation of southern New Mexico, USA. New Mexico Museum of Natural History and Science, Bulletin, 53, 52–70

Fishes and tetrapods in the Upper Pennsylvanian (Kasimovian) Cohn Coal Member of the Mattoon Formation of Illinois, United States: Systematics, Paleoecology, and Paleoenvironments. Palaios, 26(10), 639–657
DOI: 10.2110/palo.2010.p10-121r

Middle Palaeozoic microvertebrate assemblages and biogeography of East Gondwana (Australasia, Antarctica). Palaeoworld, 19(1–2), 37–54
DOI: 10.1016/j.palwor.2009.11.001
New specimens of xenacanth and hybodont sharks (Elasmobranchii: Xenacanthida and Hybodontoidea) from the Lower Permian of southwestern Oklahoma. Oklahoma Geology Notes, 63(4), 136–147

Fish Assemblages in the Tournaisian-Viséan environments of the East European Platform. In Strogen, P., Somerville, I. D. and Jones, G. L. (Eds.), Recent Advances in Lower Carboniferous Geology. Geol. Soc. Spec. Publ., 107, p. 387–415.

Vertebrate Microfossils from the Lueders Formation, Albany Group, and the faunal transition from the Wichita Group into the Clear Fork Group, Lower Permian of Texas. Modern Geology, 20, 371–382

Reworked Courceyan (Early Carboniferous) chondrichthyan remains from British Triassic cave deposits. In Turner, S.(Ed.) IGCP328: SDS The Gross Symposium, Göttingen 31 July – 7 Aug. 1993. Abstracts. 2pp.

Early Carboniferous Shark Remains from the Rockhampton District, Queensland. Memoirs of the Queensland Museum, 28(1) , 65–73

Clear Fork vertebrates and environments from the lower Permian of North-Central Texas. The Texas Journal of Science, 39(3): 253–266

Middle Palaeozoic elasmobranch remains from Australia. Journal of Vertebrate Paleontology, 2(2), 117–131
DOI: 10.1080/02724634.1982.10011923

Fossil fishes from the marine Late Palaeozoic of Holm Land-Amdrup Land, North-East Greenland. Meddelelser om Grønland, 195(9), 1–38

CHAPTER 26 Pisces In The Fossil Record. HARLAND, W.B. et al. (Eds.) 1967, London (Geological Society), pp: 627-683
Catalogue of Canadian fossil fishes. Life Sciences Contribution of the Royal Ontario Museum, 68, 1–154

Bradyodont sharks in the Permian of Western Australia. American Journal of Science, 243, 543–552

Fish remains from the Salem Limestone of Indiana. In Cumings E. R., Branson E. B., Beede, J. W. and Smith, E. A. (Eds.), The fauna of the Salem Limestone of Indiana. Rep. Indiana Dept. Geol. Natur. Resources, 30: 1376–1394

On some new species of fossil fish from the Millstone Grit rocks, with an amended list of genera and species. Proceedings of the Yorkshire Geological and Polytechnic Society, 15, 70-78
DOI: 10.1144/pygs.15.1.70
				
				
    					
    					
    					
    					
                        
    					








