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NEWSLETTER 09/2023 13.09.2023

Please acknowledge use of the database in your publications, and cite: 

Pollerspöck, J. & Straube, N. 2023, Bibliography database of living/fossil sharks, rays and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali),, World Wide Web electronic publication, Version 2023

Open Access!
We have updated our
List of Valid Extant Species
List of Described Extant Species
Free download via researchgate
The table and provided download links below are intended for informational use in Chondrichthyan research. The allocation aims for faciliating to find species numbers and most recent information on taxonomic changes. We will regularly update the table and download links at lest twice annually. The updates will be announced on facebook ( and in our monthly newsletter (sign up here: The Excel sheet allows for the application of individual filter- and sorting options. The list of described spsecies complements taxonomic information for the list of valid species by providing synonyms and / or new taxonomic combinations.
Our new paper from the team shark-references is out ????
Thanks to Iris Feichtinger and Kent Albin Nielsen for this great team work!

Jürgen Pollerspöck, Kent Albin Nielsen, Iris Feichtinger & Nicolas Straube 2023. New records of fossil deep-sea shark teeth from the Lillebælt Clay (Early–Middle Eocene) of Denmark. Bulletin of the Geological Society of Denmark, Vol. 72, 153–173. ISSN 2245-7070.

free download via researchgate

This study describes for the first time a number of distinct fossil teeth documenting several deep-sea shark species from the Eocene, which were previously not recorded from the North Sea Basin, including Apristurus sp., Orthechinorhinus cf. pfeili, Deania cf. angoumeensis, Squaliolus sp., Etmopterus cf. cahuzaci and Paraetmopterus nolfi. Our findings significantly increase the deep-sea shark diversity documented from this area so far. Despite the fact that the North Sea Basin had already lost direct connections to the neighbouring marine areas in the Eocene, the fauna shows highest similarities with documented Eocene deep-sea faunas of France, Austria and northern Morocco using cluster analysis.


We started to change our old (and unique!) citation style to adapt to the APA citation style (for information please see: to make the usage of references listed in shark references easier and more compatible with a widely accepted reference style adopted by several international scientific journals. The transition is ongoing, so far 26178 (last month: 26071) references are changed.


Since we were asked several times, if we could help distributing chondrichthyan-related job opportunities, we would like to try this out as a new category in the newsletter besides postings on our Facebook page. This category definitely depends on the community sharing job openings, so please do not hesitate and send us vacancies or similar.
Right now, we have four interesting job openings. We will keep it simple and just crosslink:


Would you like to become a shark-reference partner? Please contact us per E-mail!

Partner in Google-Maps



NEW SECTION: From now on, we will report last month’s most popular three papers from our Shark References Facebook page:

If you would like us to post information about your newly published work, please send us a picture and the paper as a pdf to or

Nr. 1 (43 Likes/Emojis, 13 Shares):

New records of the Smalltooth Sand tiger shark
Three #Odontaspis ferox were found in the English Channel and Celtic Sea in 2023. These are the first records of O. ferox in either country, and extend the species’ range by over three degrees of latitude, to >52°N. They were ~275 (female), 433 (female), and 293 cm (male) total length, respectively. These continue a series of new records, possibly indicative of a climate change induced shift in the species’ range.
Curnick, D.J. & Deaville, R. & Bortoluzzi, J.R. & Cameron, L. & Carlsson, J. & Carlsson, Je. & Dolton, H.R. & Gordon, C.A. & Hosegood, P. & Nilsson, A. & Perkins, M.W. & Purves, K.J. & Spiro, S. & Vecchiato, M. & Williams, R. & Payne, N.L. 2023 Northerly range expansion and first confirmed records of the smalltooth sand tiger shark, Odontaspis ferox, in the United Kingdom and Ireland. Journal of Fish Biology, in press
for a review of the records of the smalltooth sand tiger shark see a paper co-authord by a member of team Shark-references:
open access:
Barcelos, L.M.D. & Azeved, J.M.N. & Pollerspöck, J. & Barreiros, J.P. 2018 Review of the records of the smalltooth sand tiger shark, Odontaspis ferox (Elasmobranchii: Lamniformes: Odontaspididae), in the Azores. Acta Ichthyologica et Piscatoria, 48(2), 189–194
image: Odontaspis ferox (Risso, 1810), © Avi Klapfer, Undersea Hunter, Costa Rica

Nr. 2 (17 Likes/Emojis, 2 Shares):

open access
Chiaramonte, G.E. & Tamini, L.L. & Pettovello, A.D. & Gosztonyi, A.E. 2024 Annotated Systematic Checklist of the Ichthyofauna of the Ría Deseado (Patagonia, Argentina). Revista del Museo Argentino de Ciencias Naturales, n.s. 25(2), 207–223
We present the first complete and commented list of the ichthyofauna of the Ría Deseado (47°45´S, 65°55´W) and record the southernmost limit distribution of five species. The systematic checklist comprises 44 species belonging to 4 classes, 15 orders, 26 families, and 35 genera. The largest Families are Nototheniidae ( and Zoarcidae (6). The ichthyofauna is mainly composed of Magellanic fauna, while some Bonaerensean components are also present. Our results broaden the number of known fish species in the region, contributing as an input for the development of conservation and management plans for this area.
image: Fig. 7. Myliobatis ridens captured by Alejandro Pettovello at Caleta Tiburón, January 1996, thanks to Gustavo Enrique Chiaramonte for the paper

Nr. 3 (15 Likes/Emojis, 5 Shares):

open access!
Barbato, M. & Zampieri, C. & D'Acunto, S. & Pennino, M.G. & Barausse, A. & Mazzoldi, C. 2023 Too young to die: Mapping nursery areas for early juveniles of the endangered sandbar shark (Carcharhinus plumbeus) to inform conservation in the Mediterranean Sea. Journal of Applied Ecology, in press
Globally, elasmobranch species have been declining in abundance due to fishery activities. This conservation issue calls for effective management strategies and increasing efforts to protect these species. The declining status of elasmobranchs in the Mediterranean Sea is alarming as well. Reversing such dramatic trends requires tackling fishing pressure using multiple methods, including a fine-tuned spatial resolution in conservation strategies incorporating robust evidence on species spatial use at different life stages and its overlap with fishing pressure. In particular, a scientifically sound identification of nursery grounds is crucial to define key spatial management targets promoting the recruitment of such depleted species.
Here we focused on the sandbar shark (Carcharhinus plumbeus), listed as endangered by the IUCN, and on the nursery of its early juveniles such as newborns (i.e. a few months old) and young-of-the-year individuals (i.e., < 1 year old), both characterized by a total length below 71 cm. First, by monitoring a small-scale fishery fleet in the North-Western Adriatic Sea, we unambiguously identified a local multiyear nursery site for early juveniles of this species.
image: Carcharhinus plumbeus (Nardo, 1827), ERB 1241, female, 116, 4 TL © Frederik H. Mollen (Elasmobranch Research Belgium)


New Images

Many thanks to the following people for providing images:

Frederik H. Mollen (Elasmobranch Research Belgium) for images of  Raja straeleni Poll, 1951

Jesco Seifert for images of 
Centrophorus isodon (Chu, Meng & Liu, 1981)
Furgaleus macki (Whitley, 1943)
Centrophorus harrissoni McCulloch, 1915
Centrophorus westraliensis White, Ebert & Compagno, 2008
Centrophorus uyato (Rafinesque, 1810)


Missing papers:

Many thanks to all friends of shark-references, who sent us some missing papers last month!

Shark-References would kindly like to ask you for your contribution to this project.

At the moment we are looking for some of the following papers:

Extinct Chondrichthyes:

new entry: Priem, R. (1920) Poissons fossils du Miocène d’Egypte. Burdigalien de Moghara, „Désert libyque“. In Fourtau: Contribution à l’étude des vertébrés miocènes de l’Egypte. Cairo 1920, pp. 8-15.

new entry: Arambourg, C. & Joleaud, L. (1943) Vertébrés fossiles du basin du Niger. Bulletin Direction des Mines, 7, 1–74

new entry:  Arambourg, C. (1954) Les Poissons Crétacés du Jebel Tselfat (Maroc). Notes et Mémoires du Service Géologique du Maroc, 118: 188 pp 18 Taf.

Numano, M. (1993) Some Neogene shark-teeth from Mogami area, Yamagata Prefecture. Applied Geology of Yamagata, 13: 32–49

new entry:  Schmitz, L. (2003) Fischzähne (Neoselachii; Actinopterygii) aus dem Unter-Barremium von NW-Deutschland. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 227, 175–199

Extant Chondrichthyes:

new entry: Trois, E.F. (1877 ) Notizie sopra l'Echinorhinus spinosus osservato per la prima volta nell'Adriatico. Atti del Reale Istituto Veneto di S. L. A., Serie 5(3), 1179-1183

Chu, Y.-T.  (1930) A new species of the swallow ray (Pteroplatea) from China. China Journal, 12(6): 357

new entry: Cipria, G.  (1937) Embrione di Echinorhinus spinosus Gmelin. Memorie R Comitato Talassografico Italiano, 245, 3–7 

Smith, J.L.B. (1958) The mystery killer, the new shark Carcharhinus vanrooyeni. Veld & Vlei, 3 (9): 12–14, 28.

Deng, S.-M. & Xiong, G.-Q. & Zhan, H.-X. (1988) The deep water fishes of the east China Sea. Xue Lin Publishing house: 356 pp.

new entry: Barry, J.P. & Maher, N. (2000) Observations of the prickly shark, Echinorhinus cookei, from the oxygen minimum zone in Santa Barbara Basin, California. California Fish and Game, 86(3), 213–215

Please support and send missing papers (not listed papers or papers without the info-symbol) to or



Upcoming Meetings:


Indo-Pacific Fish Conference and the Australian Society for Fish Biology

20-24 November 2023, Auckland, New Zealand
Keep me updated

We look forward to welcoming you to the University of Auckland, located in the heart of Auckland city.  We invite you to come and meet the people that live and work here, explore our beautiful city and hope that you leave with lasting friends, partnerships and memories.

The Organising Committee look forward to welcoming you to the 11th Indo-Pacific Fish Conference (IPFC) and Annual Conference of the Australian Society for Fish Biology, to be held 20-24 November 2023 at the University of Auckland, New Zealand.

The Indo-Pacific Fish Conference (IPFC), held every four years, is undoubtedly one of the world’s premier ichthyological conferences and is eagerly anticipated by marine, estuarine and freshwater fish enthusiasts alike.

The Australian Society for Fish Biology (ASFB), founded in 1971, aims to promote research, education and management of fish and fisheries across the Indo-Pacific.

We are excited to bring these two conference together in a joint meeting that will reflect the extraordinary biological, environmental and cultural diversity of the vast Indo-Pacific region.

Auckland is a modern city offering a variety of cultural experiences, accommodation and entertainment options for every taste and budget. The city is a key regional hub, with transport connections to multiple cities across New Zealand, Australia, the Pacific and beyond.

The University of Auckland is an internationally recognised university that provides outstanding conference facilities and conference support within easy walking distance of the city centre. An exciting programme of conference field trips will allow attendees to sample the diversity of regional marine and freshwater ecosystems. Before or after the conference, delegates could explore Auckland’s magnificent Hauraki Gulf and its beautiful islands. The jewel of the gulf is Waiheke Island, a haven of vineyards, olive groves, beaches and fine dining, just a 40-minute ferry ride from downtown Auckland. For an exciting day trip, discover the history and sandy coves of Rotoroa Island, explore the open wildlife sanctuary of Tiritiri Matangi Island or climb the volcanic cone of Rangitoto Island for incredible views from the summit. If you have a little more time, travel south for a summer holiday to discover the majestic Milford and Doubtful Sounds, encounter marine life in Abel Tasman National Park, enjoy New Zealand’s beautiful Bay of Islands, or maybe plan a field trip or tropical holiday on one of the many Pacific islands that are only a short flight away from Auckland.

We look forward to welcoming you to Auckland in 2023!


The 8th edition of the
International Meeting on the Valorization and Preservation of Paleontological Heritage (RIV3P8)
November 23 - 25, 2023 (El Jadida - Morocco)

You can, also, download the 1st circular at

Would you be so kind, dear colleague, to assure a large diffusion around your colleagues paleontologists, archaeologists and prehistorians, and anyone interested in the enhancement and preservation of geoheritage (natural and cultural) from the perspective of sustainable development (see the attached poster).. Many thanks in advance. The registration is open till September 15, 2023.



To the German Ichthyological Society

Save the Date - 19th Meeting of the Society for Ichthyology (GfI) e.V. October 12-15, 2023 at the University of Applied Sciences in Bremen


Annual conference


94th Annual Meeting of the Palaeontological Society (PalGes) in Jena, 18.-22.9.2023

From Early Life to the Neandertals

For the first time since founding of the Palaeontological Society (PalGes) in Greifswald 1912 the annual meeting takes place in Jena. This is only the second time in Thuringia since the 1925 meeting in Weimar. We, from the Institute of Geosciences (IGW) at the Friedrich Schiller University of Jena, are glad to invite the members of the Society to our city at the river Saale. The city is a special location, stretching along the river, which is carving into a Muschelkalk plateau with Buntsandstein outcropping below. The surrounding Muschelkalk heights provide marvellous sights over the city. Our institute is located on the slope of the Hausberg in the eastern part of Jena. It was re-founded 1992 and still grows with an increasing number of scientists. Jena houses the most important university of Thuringia and is also a flourishing economic centre with companies like ZEISS and SCHOTT, it is popular for its touristic sights and a wide range of restaurants. The Thuringian palaeontology has a long tradition and is famous for fossils from the Permian, Triassic and Quaternary. We can offer a diverse range of excursions covering fossil sites from the late Palaeozoic to the Quaternary. The organisation of the meeting is supported by colleagues from Thüringischer Geologischer Verein (TGV), Stiftung Schloss Friedenstein Gotha, Museum Schloss Bertholdsburg Schleusingen, Research Station of Quaternary Palaeontology Weimar, and Phyletical Museum Jena. We are looking forward to welcome you in Jena and hear your presentations at 94. Annual Meeting of the PalGes in Jena!

The first circular for the meeting was published today and can be downloaded HERE.


White Sharks Global

Port Lincoln, South Australia, Australia

Sunday 12th to Friday 17th of November 2023

The organising committee is pleased to announce the upcoming White Sharks Global conference (Sunday 12th – Friday 17th of November 2023) in Port Lincoln, South Australia, home of the world’s first white shark tourism industry.
White Sharks Global is the first international white shark conference in 13 years and will provide a forum for the white shark community and stakeholders to meet, share ideas, and update information and report on recent scientific studies. This conference and associated workshops will facilitate in-depth discussions of key challenges related to white sharks.
For more information visit: and follow @WhiteSharksGlob or contact
The last white shark-focused conference was in Hawaii 13 years ago in early 2010 and a lot has research and studies have happened since. 
We have planned for five days of conference, with one day free in the middle to allow for a dive trip to the Neptune Islands Group Marine Park. There will be several focused workshops (e.g., supporting the recovery of white shark populations, supporting the management of white shark tourism, managing human-shark conflict), contributed talk sessions (5- and 15-min talks), and poster sessions. Based on previous events and our survey, we are expecting ~150 attendants from across the globe, including research scientists, students, resource managers, public safety officials, wildlife tourism operators, environmental consultants, natural history-based production companies, and television network representatives from countries all over the world such as South Africa, United States, Australia, Mexico, New Zealand, and Canada.
[On behalf of the organising committee: Charlie Huveneers (Australia), Christopher Lowe (California), Alison Towner (South Africa), Oscar Sosa-Nishizaki (Mexico), Lauren Meyer (Australia), and Greg Skomal (northwest Atlantic)]


Join us for EEA 2023, Brighton, UK!

18 - 20th October
The location is a vibrant seaside town with excellent connections to both Heathrow and Gatwick Airports and the Eurotunnel terminus in London. We will only be able to deliver an in-person EEA2023 if we get enough support by end July. Otherwise,`an online alternative will be organised.

To reserve your space at EEA 2023 please click on the link below and select "Get Tickets." You will not be charged now. We will send a link to buy tickets once a final decision is made.
Reserve EEA 2023 ticket

Abstract Submission

It’s a fast turnaround so abstract submission is open for just 6 weeks!

Please use the link below and complete this form to submit all details for your proposed presentation to the European Elasmobranch Association Conference 2023.

Please complete all sections as they should appear in the abstract book.

Submit Abstract

Extant Chondrichthyes:
Koerber, S. & Kast, S. (2023) Raja cyanoplax nom.nov., a replacement name for Raja magellanica Steindachner, 1903, junior homonym of Raja magellanica Philippi, 1902 (Chondrichthyes: Arhynchobatidae). Historia Natural (tercera serie), 13(2), 53-64
Nom nov.: Raja cyanoplax
Abstract: The homonymy between Raja magellanica Philippi, 1902 and Raja magellanica Steindachner, 1903 is removed by the introduction of a replacement name for the younger name. The discovery of a so far missing syntype in Berlin is presented, together with comments on the provenance of another syntype hosted in Vienna.  

Extinct Chondrichthyes:
no taxonomic news this month!

Rodriguez-Ibarra, E., Violante-Gonzalez, J. & Monks, S. (2023) Two new species of Acanthobothrium Blanchard, 1848 (Cestoda, Onchoproteocephalidea) from Urobatis jamaicensis (Cuvier, 1816) (Elasmobranchii, Batoidea, Myliobatiformes) of the Mexican Caribbean. Zookeys, (1169), 175–201
New species: Acanthobothrium garciaprietoi, Acanthobothrium pulidofloresae
Abstract: Eight specimens of Urobatis jamaicensis were collected from four localities of the Yu-catan Peninsula, Mexico, of which four specimens were infected with cestodes of two new species of Acanthobothrium. Acanthobothrium garciaprietoi sp. nov. differs from congeners by a combination of characters including the size of the scolex and bothridia, length of the cephalic peduncle, length of the axial and abaxial prongs and total length of the abaxial prongs of the hooks, size of the cirrus sac and testes in mature proglottids, and the total number of proglottids. The most prominent characteristic distinguishing A. pulidofloresae sp. nov. from other species of the Atlantic Ocean/Caribbean Sea/Gulf of Mexico is the form of the scolex, which has the "clover leaf" configuration. In addition, it can be distinguished by the total length of the worm, total number of proglottids, small accessory suckers, the form of the bothridia, length of the axial and abaxial prongs and total abaxial prong length of hooks, and the number of testes. According to the current category scheme, A. garciaprietoi sp. nov. and A. pulidofloresae sp. nov., belong to cat-egories 1 and 5, respectively.
Jimi, N., Shinji, J., Hookabe, N., Okanishi, M., Woo, S.P. & Nakano, T. (2023) A New Species of Branchellion (Hirudinea: Piscicolidae) Parasitizing the Gills of Short-tail Stingrays (Batoidea: Dasyatidae) From the West Pacific. Zoological Science, 40(4), 308–313
New species: Branchellion brevicaudatae
Abstract: A new fish leech, Branchellion brevicaudatae sp. n., is described based on specimens parasitizing the gills of the short-tail stingray, Bathytoshia brevicaudata (Hutton, 1875), collected from Japanese waters. The new species can be distinguished from other congeners by having: i) pulsating vesicles emerging from posterior base of branchiae, one pair per somite; ii) dorsal white spots, not arranged in longitudinal row; and iii) blackish body. A phylogenetic tree based on partial sequences of the mitochondrial cytochrome c oxidase subunit I gene from the new species and other piscicolid worms showed that the new species is sister to Branchellion torpedinis Savigny, 1822. This is the first record of Branchellion Savigny, 1822 from Japanese waters.
Bueno, V.M. & Caira, J.N. (2023) Phylogenetic relationships, host associations, and three new species of a poorly known group of "tetraphyllidean" tapeworms from elasmobranchs. Zootaxa, 5254(1), 30–50
New species: Pentaloculum grahami Zyxibothrium duffyi Zyxibothrium healyae
Abstract: This paper aims to expand understanding of a poorly known group of cestodes that parasitize an intriguingly diverse suite of elasmobranchs. The group's three currently described members (i.e., Pentaloculum macrocephalum, Pentaloculum hoi, and Zyxibothrium kamienae) parasitize an electric ray, a carpet shark, and a skate, respectively. Pentaloculum grahami n. sp. is described from a second genus of carpet shark, specifically Parascyllium collare, in Australia. Zyxibothrium duffyi n. sp. and Zyxibothrium healyae n. sp. are described from the deep-sea skates Brochiraja asperula and Brochiraja spinifera, respectively off New Zealand. The three new species share distinctive bothridia that bear a small number of large, circular, facial loculi and lateral bands of vitelline follicles that converge posterior to the ovary-features which are found in all other members of these genera. Zyxibothrium healyae n. sp. is unique in possessing three, rather than four or five, facial loculi. Zyxibothrium duffyi n. sp. possesses a combination of five facial loculi and vitelline follicles that stop short of the anterior margin of the proglottid. Pentaloculum grahami n. sp. is the largest member of the group with the greatest number of proglottids. Based on striking similarities in scolex morphology, Pentaloculum and Zyxibothrium have been hypothesized to belong to a distinct subgroup of "tetraphyllideans" provisionally designated as Clade 1. Based on sequence data for the D1-D3 region of the 28S rDNA gene generated for species of Zyxibothrium for the first time, we confirm the reciprocal monophyly of both genera as well as the monophyly of Clade 1 and its status as a distinct lineage among the "Tetraphyllidea". This work also suggests that the presence of five facial loculi is homoplasious given this character state is found in members of both genera. The new species expand the host associations of Clade 1 to include additional skate and carpet shark genera. Moving forward we would expect to find additional members of this group parasitizing other species of parascyliid carpet sharks as well as other species of the rajid genus Malacoraja and the arhynchobatid genus Brochiraja. Here we have doubled the number of described species in the taxon referred to as Clade 1 while simultaneously expanding our understanding of the morphology and anatomy of its members. This additional information will help inform the ultimate revision of the ordinal classification of the cestodes to address the highly polyphyletic nature of the order "Tetraphyllidea" as it is currently configured.


PLEASE send your new papers to or   

Latest Research Articles

Extant Chondrichthyes:
Adachi, A. & Roque, P.C.G. & Hazin, F.H.V. & Vianna, M. & Rotundo, M.M. & Oliveira, C. & Foresti, F. & Cruz, V.P. (2023): Genetic Population Structure and Diversity of the Whitetail Dogfish Squalus albicaudus (Chondrichthyes, Squaliformes) along the Brazilian Coast as Identified by SNP Markers. Fishes, 8(7), Article 373
Afonso, A.S. & Fidelis, L. (2023): The fate of plastic-wearing sharks: Entanglement of an iconic top predator in marine debris. Marine Pollution Bulletin, 194, Article 115326
Aguilar, L.A.B. & Field, C.L. & Delaune, A.J. & Aflalo, R.A. & Atkins, A. & Clauss, T.M. (2023): Retrospective review of propofol anesthesia in multiple elasmobranch species at Georgia Aquarium, 2010-2022. Journal of Zoo and Wildlife Medicine, 54(2), 326–331
Araujo, C. & Carneiro, P. & Fidelis, L. & Nascimento, B. & Antunes, M. & Viana, D. & Oliveria, P. & Torres, R. & Hazin, F. & Adam, M. (2023): Comparative genomic damage among three shark species with different habits: Sublethal impacts of human origin in a protected island environment in the South Atlantic. Marine Pollution Bulletin, 191, Article 114924
Arnette, S. & Saffarian, J. & Ferry, L. & Farina, S. (2023): Effect of parabranchial position on ventilatory pressures in the Pacific spiny dogfish (Squalus suckleyi). Zoology, 159, Article 126106
Barbato, M. & Zampieri, C. & D'Acunto, S. & Pennino, M.G. & Barausse, A. & Mazzoldi, C. (2023): Too young to die: Mapping nursery areas for early juveniles of the endangered sandbar shark (Carcharhinus plumbeus) to inform conservation in the Mediterranean Sea. Journal of Applied Ecology, in press
Barry, C. & Legaspi, C. & Clarke, T.M. & Araujo, G. & Bradshaw, C.J.A. & Gleiss, A.C. & Meyer, L. & Huveneers, C. (2023): Estimating the energetic cost of whale shark tourism. Biological Conservation, 284, Article 110164
Bellodi, A. & Mulas, A. & Daniel, L. & Cau, A. & Porcu, C. & Carbonara, P. & Follesa, M.C. (2023): Ontogenetic Shifts in Body Morphology of Demersal Sharks’ Species (Order: Squaliformes) Inhabiting the Western-Central Mediterranean Sea, with Implications for Their Bio-Ecological Role. Biology, 12(8), Article 1150
Bettcher, V.B. & Franco, A.C.S. & dos Santos, L.N. (2023): Habitat-use of the vulnerable Atlantic Nurse Shark: a review. Peerj, 11, Article e15540
Bielmyer-Fraser, G.K. & Franks, B. & Somerville, R. & Hueter, R. & Newton, A.L. & Fischer, C. (2023): Tissue metal concentrations and antioxidant enzyme activity in western north Atlantic white sharks (Carcharodon carcharias). Aquatic Toxicology, 261, Article 106641
Bizzarro, J.J. & Dewitt, L. & Wells, B.K. & Curtis, K.A. & Santora, J.A. & Field, J.C. (2023): A multi-predator trophic database for the California Current Large Marine Ecosystem. Scientific Data, 10(1), Article 496
Bouyoucos, I.A. & Shaughnessy, C.A. & Anderson, W.G. & Dores, R.M. (2023): Molecular and pharmacological analysis of the melanocortin-2 receptor and its accessory proteins Mrap1 and Mrap2 in a Squalomorph shark, the Pacific spiny dogfish. General and Comparative Endocrinology, 342, Article 114342
Bowlby, H.D. & Dicken, M.L. & Towner, A.V. & Waries, S. & Rogers, T. & Kock, A. (2023): Decline or shifting distribution? A first regional trend assessment for white sharks (Carcharodon carcharias) in South Africa. Ecological Indicators, 154, Article 110720
Bravo-Ormaza, E. & Arauz, R. & Bessudo, S. & Hearn, A. & Klimley, A.P. & Ladino-Archila, F. & Lopez-Macias, J. & Steiner, T. & Penaherrera-Palma, C. (2023): Scalloped hammerhead shark Sphyrna lewini relative abundance comparison in three offshore marine protected areas of the Eastern Tropical Pacific. Environmental Biology of Fishes, 106(8), 1767–1784
Brodbeck, B.E. & Lyons, K. & Miller, N.R. & Mohan, J.A. (2023): Sex influences elemental variation in the mineralized vertebrae cartilage of round stingray (Urobatis halleri). Marine Biology, 170(10), Article 117
Calle-Mor, M.D. & Loor-Andrade, P.J. & Galv, F. (2023): Trophic chain of the pelagic thresher, Alopias pelagicus, in the Southeastern Pacific Ocean: An approach through stable isotopes of carbon and nitrogen. Food Webs, 36, Article e00294
Caves, E.M. & Sutton, T.T. & Warrant, E.J. & Johnsen, S. (2023): Measures and models of visual acuity in epipelagic and mesopelagic teleosts and elasmobranchs. Journal of Comparative Physiology A: Neuroethology Sensory Neural and Behavioral Physiology, in press
Cetkovic, I. & Pesic, A. & Joksimovic, A. & Aloric, I. & Jovanovic, D. (2023): First documented record of neonate angular roughshark Oxynotus centrina from the eastern Adriatic Sea. Acta Adriatica, 64(1), 93–97
Chan, K.H. & Gowidjaja, J.A.P. & Urera, M.Q. & Wainwright, B.J. (2023): Analysis of Toxic Metals Found in Shark Fins Collected from a Global Trade Hub. Environmental Science & Technology, in press
Chiaramonte, G.E. & Tamini, L.L. & Pettovello, A.D. & Gosztonyi, A.E. (2023): Annotated Systematic Checklist of the Ichthyofauna of the Ría Deseado (Patagonia, Argentina). Revista del Museo Argentino de Ciencias Naturales, n.s. 25(2), 207–223
Collareta, A. & Farina, S. (2023): Did the smalleye hammerhead ever inhabit the Mediterranean Sea? A reappraisal of the only Italian record of Sphyrna tudes (Valenciennes, 1822). Acta Adriatica, 64(1), 45–51
Curnick, D.J. & Deaville, R. & Bortoluzzi, J.R. & Cameron, L. & Carlsson, J. & Carlsson, Je. & Dolton, H.R. & Gordon, C.A. & Hosegood, P. & Nilsson, A. & Perkins, M.W. & Purves, K.J. & Spiro, S. & Vecchiato, M. & Williams, R. & Payne, N.L. (2023): Northerly range expansion and first confirmed records of the smalltooth sand tiger shark, Odontaspis ferox, in the United Kingdom and Ireland. Journal of Fish Biology, in press
Dia, M. & El Vally, Y. & Meissa, B. & Abdallahi, I.C. & Diop, M. & Bouzouma, M.M. & Sarr, O. & Beibou, E. & Braham, C.B. & Jabado, R.W. (2023): Evolution of catches and specific composition of elasmobranchs in Mauritanian artisanal, coastal and offshore fisheries. Fisheries Research, 267, Article 106810
Doane, M.P. & Reed, M.B. & McKerral, J. & Lima, L.F.O. & Morris, M. & Goodman, A.Z. & Johri, S. & Papudeshi, B. & Dillon, T. & Turnlund, A.C. & Peterson, M. & Mora, M. & Venegas, R.D. & Pillans, R. & Rohner, C.A. & Pierce, S.J. & Legaspi, C.G. & Araujo, G. & Ramirez-Macias, D. & Edwards, R.A. & Dinsdale, E.A. (2023): Emergent community architecture despite distinct diversity in the global whale shark (Rhincodon typus) epidermal microbiome. Scientific Reports, 13, Article 12747
Dunn, N. & Curnick, D.J. & Carbone, C. & Carlisle, A.B. & Chapple, T.K. & Dowell, R. & Ferretti, F. & Jacoby, D.M.P. & Schallert, R.J. & Steyaert, M. & Tickler, D.M. & Williamson, M.J. & Block, B.A. & Savolainen, V. (2023): Environmental DNA helps reveal reef shark distribution across a remote archipelago. Ecological Indicators, 154, Article 110718
Dutton, J. & Hobbs, J.C. & Joung, S.J. & Schmidt, J.V. (2023): Mercury Concentrations in Whale Shark (Rhincodon typus) Embryo Muscle Tissue. Bulletin of Environmental Contamination and Toxicology, 111(2), Article 23
Freedman, R.M. & Anderson, J.M. & Caldow, C. & Stirling, B. & Rex, P. & Spurgeon, E. & McCullough, S. & Lyons, K. & May, J. & White, C.F. & Logan, R.K. & Meese, E. & Burns, E.S. & Clevenstine, A.J. & O' Sullivan, J. & Winkler, C. & Duncan, L. & Cajandig, M. & Lowe, C.G. (2023): Evidence of increasing juvenile white sharks' (Carcharodon carcharias) habitat use at the Northern Channel Islands. Journal of Fish Biology, in press
Garcia-Ruiz, C. & Delgado, M. & Hidalgo, M. (2022): Patterns of spatial changes in demersal species in the Gulf of Cadiz and northern Alboran Sea. Mediterranean Marine Science, 23(1), 55–68
Geoghegan, P. & Murphy, M.D. & Wilbur, A.R. (2023): The Structures of the Demersal Fish Communities of New Bedford and Gloucester Harbors, Two Massachusetts Urban Estuaries. Northeastern Naturalist, 30(2), 186–211
Giareta, E.P. & Hauser-Davis, R.A. & Abilhoa, V. & Wosnick, N. (2023): Carbonic anhydrase in elasmobranchs and current climate change scenario implications. Comparative Biochemistry and Physiology – Part A, Molecular & Integrative Physiology, 281, Article 111435
Gotama, R. & Stean, S.J. & Sparks, L.D. & Prasetijo, R. & Sebastian, P. (2023): Citizen science approach for monitoring fish and megafauna assemblages in a remote Marine Protected Area. Regional Studies in Marine Science, 64, Article 103058
Grech, D. & Asciutto, E. & Bakiu, R. & Battaglia, P. & Ben-Grira, C. & Camlik, O.Y. & Cappuccinelli, R. & Carmona, L. & Chebaane, S. & Crocetta, F. & Desiderato, A. & Domenichetti, F. & Dulcic, J. & Fasciglione, P. & Galil, B.S. & Galiya, M.Y. & Hoffman, R. & Langeneck, J. & Lipej, L. & Madrenas, E. & Martinelli, M. & Martin-Hervas, M.D. & Masala, C. & Mastrototaro, F. & Mavric, B. & Montesanto, F. & Mucciolo, S. & Othman, R.M. & Sempere-Valverde, J. & Soldo, A. & Spinelli, A. & Taskin, E. & Tiralongo, F. & Toso, A. & Trainito, E. & Trkov, D. & Vitale, D. & Zacchetti, L. (2023): New records of rarely reported species in the Mediterranean Sea (July 2023). Mediterranean Marine Science, 24(2), 392–418
Heinrich, D.D.U. & Dhellemmes, F. & Brown, C. & Huveneers, C. (2023): No evidence of time-place learning in juvenile lemon sharks, Negaprion brevirostris. Animal Behaviour, 203, 75–88
Hoogenboom, J.L. & Anderson, W.G. (2023): Using N-15 to determine the metabolic fate of dietary nitrogen in North Pacific spiny dogfish (Squalus acanthias suckleyi). Journal of Experimental Biology, 226(13), Article jeb244921
Hoogenboom, J.L. & Wong, M.K.S. & Hyodo, S. & Anderson, W.G. (2023): Nitrogen transporters along the intestinal spiral valve of cloudy catshark (Scyliorhinus torazame): Rhp2, Rhbg, UT. Comparative Biochemistry and Physiology – Part A, Molecular & Integrative Physiology, 280, Article 111418
Hoyos-Padilla, E.M. & Casanova-Santamaria, I. & Loria-Correa, J.C. & Sulikowski, J. (2023): The successful use of a submersible ultrasound to confirm pregnancy on free swimming bull sharks, Carcharhinus leucas, in a provisioned shark site. Frontiers in Marine Science, 10, Article 1193563
Jones, R.C. & Hall, K. & Crow, K.D. (2023): Are vertebrates constrained to two sets of paired appendages? The morphology, development, and evolution of pre-pelvic claspers in the Holocephali. Journal of Morphology, 284(9), Article e21632
Koerber, S. & Kast, S. (2023): Raja cyanoplax nom.nov., a replacement name for Raja magellanica Steindachner, 1903, junior homonym of Raja magellanica Philippi, 1902 (Chondrichthyes: Arhynchobatidae). Historia Natural (tercera serie), 13(2), 53-64
McCormack, J. & Karnes, M. & Haulsee, D. & Fox, D. & Kim, S.L. (2023): Shark teeth zinc isotope values document intrapopulation foraging differences related to ontogeny and sex. Communications Biology, 6(1), Article 711
Neves, J. & Giger, J.C. & Alves, V. & Soares, N. (2023): Focusing on Social Behaviors: Improving the Perceived Warmth of Sharks in an Aquarium Setting. Animals, 13(15), Article 2455
Papageorgiou, M. & Moutopoulos, D.K. (2023): Small-scale fisheries discards in the eastern Mediterranean Sea: Discarding species, quantities, practices and drivers. Fisheries Research, 267, Article 106798
Parmegiani, A. & Gobbato, J. & Seveso, D. & Galli, P. & Montano, S. (2023): First record of the bull shark Carcharhinus leucas (Valenciennes, 1839) from the Maldivian archipelago, central Indian Ocean. Journal of Fish Biology, in press
Pazzaglia, U.E. & Reguzzoni, M. & Milanese, C. & Manconi, R. & Lanteri, L. & Cubeddu, T. & Zarattini, G. & Zecca, P.A. & Raspanti, M. (2023): Skeletal calcification patterns of batoid, teleost, and mammalian models: Calcified cartilage versus bone matrix. Microscopy Research and Technique, in press
Rodenbiker, J. & Therkildsen, N.O. & Li, C.C. (2023): Global shark fins in local contexts: multi-scalar dynamics between Hong Kong markets and Mid-Atlantic fisheries. Ecology and Society, 28(3), Article 5
Saavedra-Sotelo, N.C. & Mendivil-Castro, P. & Onate-Gonzalez, E.C. (2023): Nuclear genetic structure of the white shark (Carcharodon carcharias) from the Northeastern Pacific. Latin American Journal of Aquatic Research, 51(3), 388–403
Saidi, B. & Enajjar, S. & Bradai, M.N. (2023): Vulnerability of elasmobranchs caught as bycatch in the grouper longline fishery in the Gulf of Gabes, Tunisia. Mediterranean Marine Science, 24(1), 142–155
Sauer, D.J. & Radford, C.A. & Mull, C.G. & Yopak, K.E. (2023): Quantitative assessment of inner ear variation in elasmobranchs. Scientific Reports, 13, Article 11939
Sherman, C.S. & Simpfendorfer, C.A. & Pacoureau, N. & Matsushiba, J.H. & Yan, H.F. & Walls, R.H.L. & Rigby, C.L. & VanderWright, W.J. & Jabado, R.W. & Pollom, R.A. & Carlson, J.K. & Charvet, P. & Bin Ali, A. & Fahmi, Cheok, J. & Derrick, D.H. & Herman, K.B. & Finucci, B. & Eddy, T.D. & Palomares, M.L.D. & Avalos-Castillo, C.G. & Kinattumkara, B. & Blanco-Parra, M. & Dharmadi, Espinoza, M. & Fernando, D. & Haque, A.B. & Mejia-Falla, P.A. & Navia, A.F. & Perez-Jimenez, J.C. & Utzurrum, J. & Yuneni, R.R. & Dulvy, N.K. (2023): Half a century of rising extinction risk of coral reef sharks and rays. Nature Communications, 14(1), Article 15
Smoothey, A.F. & Niella, Y. & Brand, C. & Peddemors, V.M. & Butcher, P.A. (2023): Bull Shark (Carcharhinus leucas) Occurrence along Beaches of South-Eastern Australia: Understanding Where, When and Why. Biology, 12(9), Article 1189
Soares, K.D.A. & Petean, F.F. (2023): Three decades of Chondrichthyan research in Brazil assessed from conferences' abstracts: patterns, gaps, and expectations. Neotropical Ichthyology, 21(2), Article e230027
Svarachorn, T. & Temple, A.J. & Berggren, P. (2023): Marine megafauna catch in Thai small-scale fisheries. Aquatic Conservation-Marine and Freshwater Ecosystems, in press
Swift, D.G. & O'Leary, S.J. & Grubbs, R.D. & Frazier, B.S. & Fields, A.T. & Gardiner, J.M. & Drymon, J.M. & Bethea, D.M. & Wiley, T.R. & Portnoy, D.S. (2023): Philopatry influences the genetic population structure of the blacktip shark (Carcharhinus limbatus) at multiple spatial scales. Molecular Ecology, in press
Tickler, D.M. & Carlisle, A.B. & Meeuwig, J.J. & Chapple, T.K. & Curnick, D.J. & Dale, J.J. & Castleton, M.J. & Schallert, R.J. & Block, B.A. (2023): Pop-up archival tags reveal environmental influences on the vertical movements of silvertip sharks Carcharhinus albimarginatus. Marine Ecology Progress Series, 717, 85–105
Torres, Y. & Charvet, M. & Faria, V.V. & Charvet, P. (2023): Dots in the dark: dorsal polychromatism in the endemic Xingu Freshwater Stingray. Journal of Zoology, in press
van der Heever, G.M. & Yemane, D. & Leslie, R.W. & van der Lingen, C.D. & Gibbons, M.J. (2023): Modeling and comparing the distributions and associations of two co-occurring catshark species off South Africa. Journal of Fish Biology, in press
van Staden, M. & Ebert, D.A. & Gennari, E. & Leslie, R.W. & McCord, M.E. & Parkinson, M. & Watson, R.G.A. & Wintner, S. & da Silva, C. & van der Merwe, A.B. (2023): Molecular Taxonomy of South Africa's Catsharks: How Far Have We Come? Diversity, 15(7), Article 828
Vazquez, D.M. & Lucifora, L.O. (2023): Estimating intrinsic susceptibility to extinction when little ecological information is available: The case of Neotropical freshwater stingrays (Chondrichthyes: Potamotrygoninae). Fish and Fisheries, in press
Vigneault, A. & Mylniczenko, N.D. & Arnold, R.D. & Jasper-Trotter, S.L. & Summa, N. & Rousseau, C. & Vergneau-Grosset, C. (2023): Management of suspected dilated cardiomyopathy with pimobendan in two leopard sharks (Triakis semifasciata). Journal of Zoo and Wildlife Medicine, 54(2), 401–405
Wagner, C.I. & Kopp, M.E.L. & Thorburn, J. & Jones, C.S. & Hoarau, G. & Noble, L.R. (2023): Characteristics of the spiny dogfish (Squalus acanthias) nuclear genome. G3-Genes Genomes Genetics, in press
Weidner, T. & Cotton, C.F. & Schieber, J.J. & Collatos, C. & Kerstetter, D.W. (2023): Short-term habitat use and vertical movements of the pelagic stingray Pteroplatytrygon violacea in the western North Atlantic Ocean determined by pop-up archival satellite tags. Bulletin of Marine Science, 99(3), 169–183
Winton, M.V. & Fay, G. & Skomal, G.B. (2023): An open spatial capture-recapture framework for estimating the abundance and seasonal dynamics of white sharks at aggregation sites. Marine Ecology Progress Series, 715, 1–25
Wosnick, N. & Charvet, P. & Hauser-Davis, R.A. & Rincon, G. & Nunes, A. & Nunes, J.L.S. (2023): Unveiling the Threats Beneath: Fish Mislabeling in the Brazilian Amazon Coast and its Impacts on the Critically Endangered Daggernose Shark. Fisheries, in press
Zimm, R. & Berio, F. & Debiais-Thibaud, M. & Goudemand, N. (2023): A shark-inspired general model of tooth morphogenesis unveils developmental asymmetries in phenotype transitions. Proceedings of the National Academy of Sciences of the United States of America, 120(15), Article e2216959120

Extinct Chondrichthyes:
Batchelor, T.J. & Duffin, C.J. (2023): A marine microvertebrate fauna from a temporary exposure of the Atherfield Clay Formation, Reigate, Surrey. Proceedings of the Geologists' Association, in Press
Burrow, C.J. & Murphy, M.A. & Turner, S. (2023): Late Silurian to earliest Devonian vertebrate biostratigraphy of the Birch Creek II section, Roberts Mountains, Nevada, U.S.A. PaleoBios, 40(4), 1–32
Collareta, A. & Merella, M. & Nobile, F. & Peri, E. & Bianucci, G. (2023): Alopias grandis (Leriche, 1942) from the Miocene of Italy: insights on a rare species of giant thresher shark. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen Band 309(2), 93–103
Dearden, R.P. & Herrel, A. & Pradel, A. (2023): Evidence for high-performance suction feeding in the Pennsylvanian stem-group holocephalan Iniopera. Proceedings of the National Academy of Sciences of the United States of America, 120(4), Article e2207854119
Hyks, P. & Kumpan, T. & Svobodova, A. (2023): Early Oxfordian occurrence of shark Notidanoides muensteri in Jurassic shelf lagoon deposits at Brno-Hady (Czechia). Palaeontographica, Abteilung A, Palaozoologie-Stratigraphie, in press
Rodriguez, D. & Ward, D.J. & Quezada, J.A. (2023): Paleontology and stratigraphic implications of a late Paleocene elasmobranch assemblage in Talcahuano, southcentral Chile. Andean Geology, 50(2), 217–247
Trif, N. & Codrea, V.A. & Pleș, G. & Bordeianu, M. (2023): The Priabonian fish from Leghia (Transylvanian Basin, Romania). Historical Biology , in press

Bueno, V.M. & Caira, J.N. (2023): Phylogenetic relationships, host associations, and three new species of a poorly known group of "tetraphyllidean" tapeworms from elasmobranchs. Zootaxa, 5254(1), 30–50
De Benedetto, G. & Capparucci, F. & Iaria, C. & Marino, F. & Gaglio, G. (2023): Helminths of the Bluntnose Sixgill Shark, Hexanchus griseus (Bonnaterre, 1788), from the Strait of Messina (Sicily, Southern Italy). Animals, 13(15), Article 2405
Isbert, W. & Dallares, S. & Grau, A. & Petrou, A. & Garcia-Ruiz, C. & Guijarro, B. & Jung, A. & Catanese, G. (2023): A molecular and epidemiological study of Grillotia (Cestoda: Trypanorhyncha) larval infection in Etmopterus spinax (Elasmobranchii: Squaliformes) in the Mediterranean Sea and Northeast Atlantic Ocean. Deep-Sea Research Part I-Oceanographic Research Papers, 199, Article 104102
Jimi, N. & Shinji, J. & Hookabe, N. & Okanishi, M. & Woo, S.P. & Nakano, T. (2023): A New Species of Branchellion (Hirudinea: Piscicolidae) Parasitizing the Gills of Short-tail Stingrays (Batoidea: Dasyatidae) From the West Pacific. Zoological Science, 40(4), 308–313
Rodriguez-Ibarra, E. & Violante-Gonzalez, J. & Monks, S. (2023): Two new species of Acanthobothrium Blanchard, 1848 (Cestoda, Onchoproteocephalidea) from Urobatis jamaicensis (Cuvier, 1816) (Elasmobranchii, Batoidea, Myliobatiformes) of the Mexican Caribbean. Zookeys, (1169), 175–201


Beyond Jaws

New episode of the podcast Beyond Jaws!

In this episode of the Beyond Jaws podcast, Dr. Lindsay Marshall shares her journey of becoming a shark science illustrator. Her illustrations are not only scientifically accurate but also incredibly detailed and captivating. From sharks and rays to leafy sea dragons, her artwork showcases a wide range of marine creatures. Dr. Marshall discusses how she turned her passion into a successful career and entrepreneurial venture. Tune in to learn more about her path in shark science and illustration.

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Sergio Bogan
Curador de Colecciones Científicas
Fundación Azara

Dear colleagues, we are editing the third issue of the year 2023 of the journal Historia Natural, whose publication is scheduled for December/January.
We would very much like to receive some ichthyology manuscripts for this new edition, if you have something you want to include or if you know someone who can contribute a manuscript, you can contact us at these email addresses

HISTORIA NATURAL is a peer-reviewed publication opened to national and international scientific community, interested in publishing original articles on Natural Sciences (Geology, Paleontology, Botany, Zoology, and Ecology). HISTORIA NATURAL is edited by the Fundación de Historia Natural “Félix de Azara” and the Centro de Ciencias Naturales, Ambientales y Antropológicas of Maimónides University. Of the three numbers that constitute the year volume, two are devoted to articles about geology, paleontology, botany, zoology and ecology, whereas the other one focuses on contributions about history of science and science workers, scientific collections and institutions. Works written in Spanish, Portuguese or English are accepted for publication. Historia Natural has a quarterly periodicity and is published on print and online formats.

Rare 14-ft smalltooth sand tiger shark washes up on Irish coast

Date: August 30, 2023
Source: Trinity College Dublin
Summary: Scientists believe a huge 14ft smalltooth sand tiger shark, which washed up at Kilmore Quay, Co. Wexford, earlier this year, represents the first of its species to have been found in Ireland's waters. Two other individuals of the same species also washed up on the UK coastline, suggesting this species' geographic range has shifted. The scientists believe these rare finds may offer a window into the future, with more and more species traditionally confined to more tropical waters expected to visit Irish and British waters.


New IUCN Shark News Newsletter is out!