NEWSLETTER 08/2022 15.08.2022

Please acknowledge use of the database www.shark-references.com in your publications, and cite: 

Pollerspöck, J. & Straube, N. 2022, Bibliography database of living/fossil sharks, rays and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali), www.shark-references.com, World Wide Web electronic publication, Version 2022

New open access paper co-authored by team shark-references!

Feichtinger, I. & Guinot, G. & Straube, N. & Harzhauser, M. & Auer, G. & Ćorić, S. & Kranner, M. & Schellhorn, S. & Ladwig, J. & Thies, D. & Pollerspöck, J. 2022 Revision of the Cretaceous shark Protoxynotus (Chondrichthyes, Squaliformes) and early evolution of somniosid sharks. Cretaceous Research, in press

Due to the peculiar combination of dental features characteristic for different squaliform families, the position of the Late Cretaceous genera Protoxynotus and Paraphorosoides within squaliform families has long been controversial. In this study, we revise these genera based on previously known fossil teeth and new dental material. The phylogenetic placement of Protoxynotus and Paraphorosoides among other extant and extinct squaliforms is discussed based on morphological characters combined with DNA sequence data for extant species. Our results suggest that Protoxynotus and Paraphorosoides should be included in the Somniosidae and that Paraphorosoides is a junior synonym of Protoxynotus. New dental material from the Campanian of Germany and the Maastrichtian of Austria enabled the description of a new species Protoxynotus mayrmelnhofi sp. nov. In addition, the evolution and origin of the characteristic squaliform tooth morphology are discussed, indicating that the elongated lower jaw teeth with erected cusp and distinct dignathic heterodonty of Protoxynotus represents a novel functional adaptation in its cutting-clutching type dentition among early squaliform sharks. Furthermore, the depositional environment of the tooth bearing horizons allows for an interpretation of the preferred habitat of this extinct dogfish shark, which exclusively occupied shelf environments of the Boreal- and northern Tethyan realms during the Late Cretaceous.

New open access paper co-authored by Dr. Straube, team shark-references!

Villate-Moreno M, Cubillos-M JC, Stibor H, Crawford AJ, Straube N.
Molecular identification and first demographic insights of sharks based on artisanal fisheries bycatch in the Pacific Coast of Colombia: implications for conservation. PeerJ 10:e13478 https://doi.org/10.7717/peerj.13478

The Pacific coast of Colombia is characterized by mangrove ecosystems which play a crucial role as possible nurseries for juvenile sharks. However, trophic food webs from coastal ecosystems are heavily disturbed by increased fishing pressure, which affects numerous shark species. In this region of the Eastern Tropical Pacific (ETP), fisheries’ data from coastal areas are scarce and unspecific, as most sharks from artisanal fisheries are landed decapitated and finless, making their morphological identification difficult. For the establishment and implementation of effective regional conservation and management policies, information on the diversity and population dynamics of shark species is crucial. We therefore sequenced the mitochondrial NADH2 gene of 696 samples taken from fishermen’s landings of shark’s bycatch along the Colombian north Pacific coast. We were able to identify 14 species of sharks, two of the most abundant species were Sphyrna lewini and Carcharhinus falciformis, both evaluated on IUCN the Red List of Threatened species (Critically Endangered and Vulnerable) and CITES regulated. We found low genetic diversity in the sampled area increasing the concern for both species in the region, even more considering that the majority of individuals were juveniles. Our results showed the importance of genetic markers for first population genetic insights as a complementary tool during the decision-making process in management plans. For this specific region, strategies such as the delimitation of conservation priority areas or the regulation of fishing gears could help improve the sustainability of shark populations in the Colombian Pacific.



We started to change our old (and unique!) citation style to adapt to the APA citation style (for information please see: https://www.mendeley.com/guides/apa-citation-guide) to make the usage of references listed in shark references easier and more compatible with a widely accepted reference style adopted by several international scientific journals. The transition is ongoing, so far 24783 (last month: 24643) references are changed.



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NEW SECTION: From now on, we will report last month’s most popular three papers from our Shark References Facebook page:

If you would like us to post information about your newly published work, please send us a picture and the paper as a pdf to nicolas.straube@shark-references.com or juergen.pollerspoeck@shark-references.com.

Nr. 1 (8.038 People reached):

Villate-Moreno M, Cubillos-M JC, Stibor H, Crawford AJ, Straube N.
Molecular identification and first demographic insights of sharks based on artisanal fisheries bycatch in the Pacific Coast of Colombia: implications for conservation.

The Pacific coast of Colombia is characterized by mangrove ecosystems which play a crucial role as possible nurseries for juvenile sharks. However, trophic food webs from coastal ecosystems are heavily disturbed by increased fishing pressure, which affects numerous shark species. In this region of the Eastern Tropical Pacific (ETP), fisheries’ data from coastal areas are scarce and unspecific, as most sharks from artisanal fisheries are landed decapitated and finless, making their morphological identification difficult. For the establishment and implementation of effective regional conservation and management policies, information on the diversity and population dynamics of shark species is crucial. We therefore sequenced the mitochondrial NADH2 gene of 696 samples taken from fishermen’s landings of shark’s bycatch along the Colombian north Pacific coast. We were able to identify 14 species of sharks, two of the most abundant species were Sphyrna lewini and Carcharhinus falciformis, both evaluated on IUCN the Red List of Threatened species (Critically Endangered and Vulnerable) and CITES regulated. We found low genetic diversity in the sampled area increasing the concern for both species in the region, even more considering that the majority of individuals were juveniles. Our results showed the importance of genetic markers for first population genetic insights as a complementary tool during the decision-making process in management plans. For this specific region, strategies such as the delimitation of conservation priority areas or the regulation of fishing gears could help improve the sustainability of shark populations in the Colombian Pacific.

Villate-Moreno M, Cubillos-M JC, Stibor H, Crawford AJ, Straube N. 2022. Molecular identification and first demographic insights of sharks based on artisanal fisheries bycatch in the Pacific Coast of Colombia: implications for conservation. PeerJ 10:e13478 https://doi.org/10.7717/peerj.13478

Shark references: https://shark-references.com/species/view/Sphyrna-lewini

Nr. 2 (7.624 People reached):


Thanks to María Cristina Oddone for this new paper!

The aim of the present study was to analyze the morphological pattern of dermal denticles across the body of the poorly known deep-water catshark Galeus mincaronei, which is endemic to Southern Brazil. Additionally, we aimed to identify differences between sexes and stages of sexual maturity. For this purpose, we analyzed patches of eight skin samples from body regions, that is, the head, body, and tail. Dermal denticles were imaged using scanning electron microscopy.

Macías‐Cuyare, M. & Oddone, M.C. 2022 Morphological pattern of the dermal denticles of the Southern sawtail catshark Galeus mincaronei Soto, 2001. Journal of Morphology, in press
Morphological pattern of the dermal denticles of the Southern sawtail catshark Galeus mincaronei Soto, 2001.

images: Scanning electron micrographs of dermal denticles from the head region, (a) N = nostril, (b) H1 = snout, and (c) H2 = cranium for developing and mature males (upper panel) and developing, immature, and mature females (lower) of the Southern sawtail catshark Galeus mincaronei in southern Brazil (×100). (→) Ampullae of Lorenzini

Nr. 3 (2.085 People reached):

Thanks to Alexei Orlov for this new open access paper!

Ontogenic Changes in Coloration of Rare Deepwater Richardson’s Ray Bathyraja richardsoni (Arhynchobatidae, Rajiformes, Chondrichthyes)

The features of the coloration of the dorsal and ventral surfaces of the rare deep-sea Richardson’s ray Bathyraja richardsoni (Arhynchobatidae) in the length range from 14 to 146 cm have been studied and its ontogenetic changes have been traced. New data on the coloration of embryos, immature, maturing, and sexually mature individuals are presented. It is shown that variations in the coloration of the dorsal surface are insignificant, while the coloration of the ventral surface can vary significantly.

image by the authors


New Images

Many thanks to the following people for providing images:

Frederik Mollen, Elasmobranch Research, Belgium for images of Orectolobus hutchinsi Last, Chidlow & Compagno, 2006

Jesco Seifert,  for images of Centrophorus moluccensis Bleeker, 1860

Sebastian Wilfling for an images of Carcharhinus longimanus (Poey, 1861)

Rodolfo Abrams courtesy of Conservación ConCiencia for a image of Scyliorhinus torrei Howell Rivero, 1936


Missing papers:

Many thanks to all friends of shark-references, who sent us some missing papers last month!

Shark-References would kindly like to ask you for your contribution to this project.

At the moment we are looking for some of the following papers:

Extinct Chondrichthyes:

Numano, M. (1993) Some Neogene shark-teeth from Mogami area, Yamagata Prefecture. Applied Geology of Yamagata, 13: 32–49

Extant Chondrichthyes:

Marini, T.L.  (1935) Nota sobre una raya argentina. Physis, 11(40): 503–506

SMITH, J.L.B. (1958) The mystery killer, the new shark Carcharhinus vanrooyeni. Veld & Vlei, 3 (9): 12–14, 28.

GUBANOV, E.P. & SCHLEIB, N.A. (1980) Sharks of the Arabian Gulf. Kuwait Ministry of Public Works, Agracultural Department, Fisheries Division. Sharks of the Arabian Gulf.: 1–69

DOLGANOV, V.N. (1983) Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod. [Manual for identification of cartilaginous fishes of Far East seas of USSR and adjacent waters.] TINRO, Vladivostok. Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod.: 92 pp.

Please support www.shark-references.com and send missing papers (not listed papers or papers without the info-symbol) to juergen.pollerspoeck@shark-references.com or nicolas.straube@shark-references.com



Upcoming Meetings:

11th International Cretaceous Symposium Warsaw, Poland, 2022

August 22-26

Abstract submission deadline
30th April 2022

Early Bird Fees 
15th May 2022

3rd circular with detailed programme 
c. 15th July 2022

Field trip registration deadline  
31st May 2022


5th International Marine ProtectedAreas Congress (IMPAC5)
September 1 – 8, 2022 Vancouver, Canada
International Marine Protected Areas Congresses (IMPAC) are an opportunity for the global community of marine conservation managers and practitioners to exchange knowledge, experience and best practices to strengthen the conservation of marine biodiversity and to protect the natural and cultural heritage of the ocean. IMPAC5 will be jointly hosted by the Host First Nations — Musqueam Indian Band, Squamish Nation, and Tsleil-waututh Nation — together with the Province of British Columbia, the Government of Canada, the Canadian Parks and Wilderness Society (CPAWS) and the International Union for the Conservation of Nature (IUCN). IMPAC5 is an opportunity to bring together Indigenous peoples and cultures from around the world to embrace a collaborative approach and learn from Indigenous leadership in ocean conservation. Join thousands of marine protected area professionals from around the world to chart a course towards protecting 30% of the ocean by 2030. Learn about traditional marine protection practices and innovative sustainability initiatives from local and international indigenous experts.


Sharks International Conference 2022 (SI2022)
October 10 – 14, 2022 (online virtual conference)
October 20 – 22, 2022 (physical in-person conference)
Valencia, Spain
SI2022 is a hybrid event in October 2022 that will bring together a strong community of people from across the world interested in sharks and rays, all in the name of addressing the challenge of elasmobranch conservation in this Decade of Ocean Science. In association with the European Elasmobranch Association (EEA) and hosted by the Shark Trust, Submon, and Lamna, the event will include five online days (October 10- 14th) featuring enhanced digital content on key themes in shark conservation, leading up to a three-day physical conference in Valencia (October 20th-22nd). The conference is funded by the Save our Seas Foundation and will be based out of L’Oceanogràfic, the largest aquarium in Europe, and streamed live across the world. If you are interested in sharks and rays and want to be a part of the 300+ member community currently shaping SI2022, sign up to the portal at si2022.org. Join this year to be automatically entitled to a 10% discount when tickets become available.




Extant Chondrichthyes:
no taxonomic news this month

Extinct Chondrichthyes:
no taxonomic news this month

Franzese, S. & Mutti, L.D. & Tropea, C. & Ivanov, V.A. (2022): Morphological study of members of the genus Echeneibothrium (Cestoda: Rhinebothriidea: Echeneibothriidae) from rajiform skates of the Argentine Sea and analysis of the phylogenetic relationships within the family Echeneibothriidae. Zoologischer Anzeiger, 299, 1–10
New species: Echeneibothrium cristinae
Abstract: A new cestode species of the genus Echeneibothrium van Beneden, 1850 was collected from a rajiform skate in the Argentine Sea: Echeneibothrium cristinae sp. nov. parasitizing Bathyraja cousseauae Díaz de Astarloa & Mabragaña, 2004, from coastal waters off Tierra del Fuego Province. This species differs from congeners by a particular combination of the following features: total length, number of proglottids, number of loculi and presence of a medial longitudinal septum on the distal bothridial surface, length of bothridial stalks, size of the cirrus sac, position of the ovarian isthmus and course of the vagina. In addition, two known Echeneibothrium species, originally reported in Dipturus chilensis (Guichenot, 1848) from the Chilean Sea, were collected from a new host in the Argentine Sea: Echeneibothrium multiloculatum Carvajal & Dailey, 1975 and Echeneibothrium williamsi Carvajal & Dailey, 1975, parasitizing the rajiform skate Dipturus brevicaudatus (Marini, 1933) from coastal waters off Buenos Aires Province (E. multiloculatum) and Patagonia (both species). Emendations were made to the descriptions and redescriptions of these species, including the modification of measurement ranges for several characters and the addition of important morphological information omitted in previous studies. Finally, a phylogenetic analysis based on morphological characters was performed to determine the inter-relationships within the family Echeneibothriidae, this study being the first to consider representatives of all known genera belonging to that family.


PLEASE send your new papers to
juergen.pollerspoeck@shark-references.com or 

Latest Research Articles

Extant Chondrichthyes:
Adams, L. & Lyons, K. & Larkin, E. & Leier, N. & Monday, J. & Plante, C. & Dubach, J. & Wyffels, J. (2022) Artificial Insemination and Parthenogenesis in the Zebra Shark Stegostoma tigrinum. Frontiers in Marine Science, 9, Article 886616 https://dx.doi.org/10.3389/fmars.2022.886616
Afonso, A.S. & Macena, B.C.L. & Mourato, B. & Bezerra, N.P.A. & Mendonca, S. & de Queiroz, J. & Hazin, F.H.V. (2022) Trophic-Mediated Pelagic Habitat Structuring and Partitioning by Sympatric Elasmobranchs. Frontiers in Marine Science, 9, Article 779047 https://dx.doi.org/10.3389/fmars.2022.779047
Afonso, P. & Gandra, M. & Graca, G. & Macena, B. & Vandeperre, F. & Fontes, J. (2022) The Multi-Annual Residency of Juvenile Smooth Hammerhead Shark in an Oceanic Island Nursery. Frontiers in Marine Science, 9, Article 844893 https://dx.doi.org/10.3389/fmars.2022.844893
Anderson, J.M. & Spurgeon, E. & Stirling, B.S. & May, J. & Rex, P.T. & Hyla, B. & McCullough, S. & Thompson, M. & Lowe, C.G. (2022) High resolution acoustic telemetry reveals swim speeds and inferred field metabolic rates in juvenile white sharks (Carcharodon carcharias). Plos One, 17(6), Article e0268914 https://dx.doi.org/10.1371/journal.pone.0268914
Ariza, A.A. & Adachi, A. & Roque, P. & Hazin, F.H.V. & Vianna, M. & Rotundo, M.M. & Delpiani, S.M. & de Astarloa, J.M.D. & Delpiani, G. & Oliveira, C. & Foresti, F. & Cruz, V.P. (2022) DNA Barcoding and Species Delimitation for Dogfish Sharks Belonging to the Squalus Genus (Squaliformes: Squalidae). Diversity-Basel, 14(7), Article 544 https://dx.doi.org/10.3390/d14070544
Augustine, S. & Lika, K. & Kooijman, S. (2022) The comparative energetics of the chondrichthyans reveals universal links between respiration, reproduction and lifespan. Journal of Sea Research, 185, Article 102228 https://dx.doi.org/10.1016/j.seares.2022.102228
Bakke, F.K. & Gundappa, M.K. & Matz, H. & Stead, D.A. & Macqueen, D.J. & Dooley, H. (2022) Exploration of the Nurse Shark (Ginglymostoma cirratum) Plasma Immunoproteome Using High-Resolution LC-MS/MS. Frontiers in Immunology, 13, Article 873390 https://dx.doi.org/10.3389/fimmu.2022.873390
Barker, J. & Davies, J. & Goralczyk, M. & Patel, S. & O'Connor, J. & Evans, J. & Sharp, R. & Gollock, M. & Wood, F.R. & Rosindell, J. & Bartlett, C. & Garner, B.J. & Jones, D. & Quigley, D. & Wray, B. (2022) The distribution, ecology and predicted habitat use of the Critically Endangered angelshark (Squatina squatina) in coastal waters of Wales and the central Irish Sea. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.15133
Blanco-Parra, M.D. & Gasca, A.A. & Rincon, C.A.R. & Martinez, N.H.G. & Nino-Torres, C.A. (2022) Citizen Science as a Tool to Get Baseline Ecological and Biological Data on Sharks and Rays in a Data-Poor Region. Sustainability, 14(11), Article 6490 https://dx.doi.org/10.3390/su14116490
Bottaro, M. (2022) Sixth sense in the deep-sea: the electrosensory system in ghost shark Chimaera monstrosa. Scientific Reports, 12(1), Article 9848 https://dx.doi.org/10.1038/s41598-022-14076-2
Caceres, C. & Kiszka, J.J. & Luna-Acosta, A. & Herrera, H. & Zarza, E. & Heithaus, M.R. (2022) Predatory fish exploitation and relative abundance in a data-poor region from the Caribbean coast of Colombia, inferred from artisanal fishery interview surveys and baited remote underwater video systems. Aquatic Conservation-Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/aqc.3853
Campbell, M.J. & Rigby, C.L. (2022) A re-examination of the growth of the gummy shark (Mustelus antarcticus) from Queensland, Australia. Marine and Freshwater Research, in press https://dx.doi.org/10.1071/mf22066
Canfield, S.J. & Galvan-Magana, F. & Bowen, B.W. (2022) Little Sharks in a Big World: Mitochondrial DNA Reveals Small-scale Population Structure in the California Horn Shark (Heterodontus francisci). Journal of Heredity, 113(3), 298–310 https://dx.doi.org/10.1093/jhered/esac008
Cardenosa, D. & Shea, S.K. & Zhang, H.R. & Fischer, G.A. & Simpfendorfer, C.A. & Chapman, D.D. (2022) Two thirds of species in a global shark fin trade hub are threatened with extinction: Conservation potential of international trade regulations for coastal sharks. Conservation Letters, in press https://dx.doi.org/10.1111/conl.12910
Carpenter, M. & Cullain, N. & Venables, S.K. & Tibirica, Y. & Griffiths, C. & Marshall, A.D. (2022) Evidence of Zavora Bay as a critical site for reef manta rays, Mobula alfredi, in southern Mozambique. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.15132
Cerutti-Pereyra, F. & Salinas-De-Leon, P. & Arnes-Urgelles, C. & Suarez-Moncada, J. & Espinoza, E. & Vaca, L. & Paez-Rosas, D. (2022) Using stable isotopes analysis to understand ontogenetic trophic variations of the scalloped hammerhead shark at the Galapagos Marine Reserve. Plos One, 17(6), Article e0268736 https://dx.doi.org/10.1371/journal.pone.0268736
Ćetković, I. & Pešić, A. & Ikica, Z. & Milošević, D. & Mrdak, D. (2022) Occurrence of rare and endangered elasmobranchs in by-catch of Montenegrin fisheries (South-Eastern Adriatic Sea). Cahiers de Biologie Marine, 63, 247–256  https://dx.doi.org/10.21411/CBM.A.37B316BB
Cevik, J. & Hunter-Smith, D.J. & Rozen, W.M. (2022) Infections following stingray attacks: A case series and literature review of antimicrobial resistance and treatment. Travel Medicine and Infectious Disease, 47, Article 102312 https://dx.doi.org/10.1016/j.tmaid.2022.102312
Clark-Shen, N. & Chin, A. & Arunrugstichai, S. & Labaja, J. & Mizrahi, M. & Simeon, B. & Hutchinson, N. (2022) Status of Southeast Asia's marine sharks and rays. Conservation Biology, in press https://dx.doi.org/10.1111/cobi.13962
Cortelezzi, P. & Paulet, T.G. & Olbers, J.M. & Harris, J.M. & Bernard, A.T.F. (2022) Conservation benefits of a marine protected area on South African chondrichthyans. Journal of Environmental Management, 319, Article 115691 https://dx.doi.org/10.1016/j.jenvman.2022.115691
Cousido-Rocha, M. & Cervino, S. & Alonso-Fernandez, A. & Gil, J. & Herraiz, I.G. & Rincon, M.M. & Ramos, F. & Rodriguez-Cabello, C. & Sampedro, P. & Vila, Y. & Pennino, M.G. (2022) Applying length-based assessment methods to fishery resources in the Bay of Biscay and Iberian Coast ecoregion: Stock status and parameter sensitivity. Fisheries Research, 248, Article 106197 https://dx.doi.org/10.1016/j.fishres.2021.106197
Cronin, M.R. & Croll, D.A. & Hall, M.A. & Lezama-Ochoa, N. & Lopez, J. & Murua, H. & Murua, J. & Restrepo, V. & Rojas-Perea, S. & Stewart, J.D. & Waldo, J.L. & Moreno, G. (2022) Harnessing stakeholder knowledge for the collaborative development of Mobulid bycatch mitigation strategies in tuna fisheries. Ices Journal of Marine Science, in press https://dx.doi.org/10.1093/icesjms/fsac093
Damalas, D. & Stamouli, C. & Fotiadis, N. & Kikeri, M. & Kousteni, V. & Mantopoulou-Palouka, D. (2022) The Gyaros island marine reserve: A biodiversity hotspot in the eastern Mediterranean Sea. Plos One, 17(2), Article e0262943 https://dx.doi.org/10.1371/journal.pone.0262943
Das, D. & Afonso, P. (2022) Review of the Diversity, Ecology, and Conservation of Elasmobranchs in the Azores Region, Mid-North Atlantic. Frontiers in Marine Science, 4, Article 354 https://dx.doi.org/10.3389/fmars.2017.00354
Das, D. & Gonzalez-Irusta, J.M. & Morato, T. & Fauconnet, L. & Catarino, D. & Afonso, P. & Viegas, C. & Rodrigues, L. & Menezes, G. & Rosa, A. & Pinho, M.R.R. & da Silva, H.M. & Giacomello, E. (2022) Distribution models of deep-sea elasmobranchs in the Azores, Mid-Atlantic Ridge, to inform spatial planning. Deep-Sea Research Part I-Oceanographic Research Papers, 182, Article 103707 https://dx.doi.org/10.1016/j.dsr.2022.103707
De Wysiecki, A.M. & Cortes, F. & Jaureguizar, A.J. & Barnett, A. (2022) Potential global distribution of a temperate marine coastal predator: The role of barriers and dispersal corridors on subpopulation connectivity. Limnology and Oceanography, in press https://dx.doi.org/10.1002/lno.12168
D'Iglio, C. & Albano, M. & Famulari, S. & Spano, N. & Rinelli, P. & Savoca, S. & Capillo, G. (2022) Basic Intersexuality (Abnormal Hermaphroditism) in the Blackmouth Catshark, Galeus melastomus, (Rafinesque, 1810), from the Southern Tyrrhenian Sea (Central Mediterranean Sea). Fishes, 7(3), Article 120 https://dx.doi.org/10.3390/fishes7030120
Dodd, J. & Baxter, J.M. & Donnan, D.W. & James, B.D. & Lavender, E. & McSorley, C.A. & Mogg, A.O.M. & Thorburn, J.A. (2022) First report of an egg nursery for the Critically Endangered flapper skate Dipturus intermedius (Rajiformes: Rajidae). Aquatic Conservation-Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/aqc.3857
Dolganov, V.N. & Tuponogov, V.N (2022) Identification table (key) of skates of genera Bathyraja and Rhinioraja (family Rajidae) of Far-East seas of Russia [in Russian]. Izvestiya TINRO, 126, 657–663
Du, J.G. & Ding, L.K. & Su, S.K. & Hu, W.J. & Wang, Y.Y. & Loh, K.H. & Yang, S.Y. & Chen, M.R. & Roeroe, K.A. & Songploy, S. & Liu, Z.H. & Chen, B. (2022) Setting Conservation Priorities for Marine Sharks in China and the Association of Southeast Asian Nations (ASEAN) Seas: What Are the Benefits of a 30% Conservation Target? Frontiers in Marine Science, 9, Article 933291 https://dx.doi.org/10.3389/fmars.2022.933291
Espino, F. & Gonzalez, J.A. & Bosch, N.E. & Otero-Ferrer, F.J. & Haroun, R. & Tuya, F. (2022) Distribution and population structure of the smooth-hound shark, Mustelus mustelus (Linnaeus, 1758), across an oceanic archipelago: Combining several data sources to promote conservation. Ecology and Evolution, 12(7), Article e9098 https://dx.doi.org/10.1002/ece3.9098
Fakioglu, Y.E. & Ozbilgin, H. & Gokce, G. & Herrmann, B. (2022) Effect of ground gear modification on bycatch of rays in mediterranean bottom trawl fishery. Ocean & Coastal Management, 223, Article 106134 https://dx.doi.org/10.1016/j.ocecoaman.2022.106134
Falsone, F. & Gancitano, V. & Geraci, M.L. & Sardo, G. & Scannella, D. & Serena, F. & Vitale, S. & Fiorentino, F. (2022) Assessing the Stock Dynamics of Elasmobranchii off the Southern Coast of Sicily by Using Trawl Survey Data. Fishes, 7(3), Article 136 https://dx.doi.org/10.3390/fishes7030136
Fernandez-Aguirre, E. & Galvan-Magana, F. & Sanchez-Gonzalez, A. & Gonzalez-Armas, R. & Abitia-Cardenas, L.A. & Elorriaga-Verplancken, F.R. & Villalejo-Fuerte, M.T. & Tripp-Valdez, A. & Barajas-Calderon, A.V. & Delgado-Huertas, A. & Torres-Garcia, R.Q. (2022) Changes in the feeding habits of the bat ray Myliobatis californica (Gill 1865) during climatic anomalies off the west coast of the Baja California Peninsula, Mexico. Regional Studies in Marine Science, 53, Article 102462 https://dx.doi.org/10.1016/j.rsma.2022.102462
Fetterplace, L.C. & Delgado Esteban, J.J. & Pini-Fitzsimmons, J. & Gaskell, J. & Wueringer, B.E. (2022) Evidence of sound production in wild stingrays. Ecology, in press https://dx.doi.org/10.1002/ecy.3812
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Shinder, S.L. & Cray, C. & Hammerschlag, N. & Merly, L. (2022) Serum Protein Electrophoresis Reference Intervals for Six Species of Wild-Sampled Sharks in South Florida. Integrative and Comparative Biology, in press https://dx.doi.org/10.1093/icb/icac063
Shipley, O.N. & Olin, J.A. & Whiteman, J.P. & Bethea, D.M. & Newsome, S.D. (2022) Bulk and amino acid nitrogen isotopes suggest shifting nitrogen balance of pregnant sharks across gestation. Oecologia, 199(2), 313–328 https://dx.doi.org/10.1007/s00442-022-05197-6
Simonitis, L.E. & Marshall, C.D. (2022) Microstructure of the Bonnethead Shark (Sphyrna tiburo) Olfactory Rosette. Integrative Organismal Biology, 4(1), Article obac027 https://dx.doi.org/10.1093/iob/obac027
Steele, L.M. & Okihiro, M.S. & Berlemont, R. & Dillon, J.G. & Young, K.A. & Hesami, S. & Van Sommeran, S. & Lowe, C.G. (2022) Carnobacterium maltaromaticum associated with meningoencephalitis and otitis in stranded common thresher sharks (Alopias vulpinus). Veterinary Pathology, in press https://dx.doi.org/10.1177/03009858221102600
Surapaneni, V.A. & Schindler, M. & Ziege, R. & de Faria, L.C. & Wölfer, J. & Bidan, C.M. & Mollen, F.H. & Amini, S. & Hanna, S. & Dean, M.N. (2022) Groovy and Gnarly: Surface Wrinkles as a Multifunctional Motif for Terrestrial and Marine Environments. Integrative and Comparative Biology, in press, Article icac079 https://dx.doi.org/10.1093/icb/icac079
Thiery, A.P. & Standing, A.S. & Cooper, R.L. & Fraser, G.J. (2022) An epithelial signalling centre in sharks supports homology of tooth morphogenesis in vertebrates. Elife, 11, Article e73173 https://dx.doi.org/10.7554/eLife.73173
Tiznado-Salazar, V.M. & Bojorquez-Sanchez, C. & Zamora-Garcia, O.G. & Marquez-Farias, J.F. & Paez-Osuna, F. & Berges-Tiznado, M.E. (2022) Bioaccumulation of Essential and Potentially Toxic Elements in the Muscle and Liver of the Spotted Ratfish (Hydrolagus colliei) From Deep-Sea Waters off the Northern Gulf of California. Biological Trace Element Research, in press https://dx.doi.org/10.1007/s12011-022-03330-3
Tokunaga, S. & Watanabe, Y.Y. & Kawano, M. & Kawabata, Y. (2022) Factors affecting gestation periods in elasmobranch fishes. Biology Open, 11(6), Article bio059270 https://dx.doi.org/10.1242/bio.059270
Towner, A.V. & Watson, R.G.A. & Kock, A.A. & Papastamatiou, Y. & Sturup, M. & Gennari, E. & Baker, K. & Booth, T. & Dicken, M. & Chivell, W. & Elwen, S. & Kaschke, T. & Edwards, D. & Smale, M.J. (2022) Fear at the top: killer whale predation drives white shark absence at South Africa's largest aggregation site. African Journal of Marine Science, 44(2), 139–152 https://dx.doi.org/10.2989/1814232x.2022.2066723
Villalobos-Segura, E. & Marrama, G. & Carnevale, G. & Claeson, K.M. & Underwood, C.J. & Naylor, G.J.P. & Kriwet, J. (2022) The Phylogeny of Rays and Skates (Chondrichthyes: Elasmobranchii) Based on Morphological Characters Revisited. Diversity, 14(6), Article 456 https://dx.doi.org/10.3390/d14060456
Villate-Moreno, M, & Cubillos-M, J.C. & Stibor, H. & Crawford, A.J. & Straube, N. (2022) Molecular identification and first demographic insights of sharks based on artisanal fisheries bycatch in the Pacific Coast of Colombia: implications for conservation. PeerJ, 10, Article e13478 https://dx.doi.org/10.7717/peerj.13478
Wheeler, C.R. & Kneebone, J. & Heinrich, D. & Strugnell, J.M. & Mandelman, J.W. & Rummer, J.L. (2022) Diel Rhythm and Thermal Independence of Metabolic Rate in a Benthic Shark. Journal of Biological Rhythms, in press https://dx.doi.org/10.1177/07487304221107843
White, W.T. & O'Neill, H.L. & Cleeland, J. & Lamb, T.D. (2022) Further description of the Kerguelen sandpaper skate Bathyraja irrasa (Rajiformes: Arhynchobatidae) based on additional specimens, including egg cases and embryos. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.15136
Whitehead, D.A. & Ayres, K.A. & Gayford, J.H. & Ketchum, J.T. & Galvan-Magana, F. & Christiansen, F. (2022) Aerial photogrammetry of whale sharks (Rhincodon typus) in the Bay of La Paz, using an unoccupied aerial vehicle. Marine Biology, 169(7), Article 94 https://dx.doi.org/10.1007/s00227-022-04085-0
Whitehead, D.A. & Gayford, J.H. & Hoyos, E.M. & Shorter, N.M. & Galvan-Magana, F. & Ketchum, J.T. (2022) Headline: frequency and distribution of mating wounds in a silky shark aggregation first description of a sex segregated aggregation of silky sharks (Carcharhinus falciformis) and the frequency and distribution of mating wounds off the tip of the Baja California Peninsula. Environmental Biology of Fishes, 105(7), 953–960 https://dx.doi.org/10.1007/s10641-022-01297-7
Whitehead, D.A. & Gayford, J.H. & Hoyos, E.M. & Shorter, N.M. & Galvan-Magana, F. & Ketchum, J.T. (2022) First description of a sex segregated aggregation of silky sharks (Carcharhinus falciformis) and the frequency and distribution of mating wounds off the tip of the Baja California Peninsula (Jul, 10.1007/s10641-022-01297-7, 2022). Environmental Biology of Fishes, 105(7), 961–961 https://dx.doi.org/10.1007/s10641-022-01304-x
Wong, D. & Smart, J.J. & Barrow, J. & Cleeland, J. & Yates, P. & Ziegler, P. & Rizzari, J.R. (2022) Age, growth and maturity of Southern Ocean skates (Bathyraja spp.) from the Kerguelen Plateau. Polar Biology, in press https://dx.doi.org/10.1007/s00300-022-03062-z
Wosnick, N. & Leite, R.D. & Giareta, E.P. & Morick, D. & Hauser-Davis, R.A. (2022) Unraveling Metabolite Provisioning to Offspring Through Parental Fluids: A Case Study of the Brazilian Guitarfish, Pseudobatos horkelii. Frontiers in Physiology, 13, Article 911617 https://dx.doi.org/10.3389/fphys.2022.911617
Yan, X.D. & Lapinski, M.K. & Gore, M. & Neuberger, L. & Grayson-Sneed, K. (2022) Understanding shark-related cognitive vulnerability and its role in shark diving: implications for conservation. Human Dimensions of Wildlife, in press https://dx.doi.org/10.1080/10871209.2022.2101720
Zamora-Arellano, N. & Ruelas-Inzunza, J. & Amezcua, F. & Spanopoulos-Zarco, P. & Ibarra-Valdez, A.K. (2022) Bioaccumulation of cd and hg in Muscle of Juvenile Pacific Sharpnose Shark Rhizoprionodon longurio from the SE Gulf of California. Bulletin of Environmental Contamination and Toxicology, in press https://dx.doi.org/10.1007/s00128-022-03550-2
Zhao, Y. & Jiang, C.P. & Ju, P.L. & Xiao, J.M. & Chen, M.R. (2022) Reproductive Biology of the Pacific Spadenose Shark Scoliodon macrorhynchos, a Heavily Exploited Species in the Southern Taiwan Strait. Marine and Coastal Fisheries, 14(4), Article e10216 https://dx.doi.org/10.1002/mcf2.10216
Extinct Chondrichthyes:
Bazzi, M. & Kear, B.P. & Siversson, M. (2022) Southern higher-latitude lamniform sharks track mid-Cretaceous environmental change. Gondwana Research, 103, 362–370 https://dx.doi.org/10.1016/j.gr.2021.10.012
Benites-Palomino, A. & Velez-Juarbe, J. & Altamirano-Sierra, A. & Collareta, A. & Carrillo-Briceno, J.D. & Urbina, M. (2022) Sperm whales (Physeteroidea) from the Pisco Formation, Peru, and their trophic role as fat sources for late Miocene sharks. Proceedings of the Royal Society of London, Series B, 289(1977), Article 20220774 https://dx.doi.org/10.1098/rspb.2022.0774
Benyoucef, M. & Perez-Garcia, A. & Bendella, M. & Ortega, F. & Vullo, R. & Bouchemla, I. & Ferre, B. (2022) The "mid"-Cretaceous (Lower Cenomanian) Continental Vertebrates of Gara Samani, Algeria. Sedimentological Framework and Palaeodiversity. Frontiers in Earth Science, 10, Article 927059 https://dx.doi.org/10.3389/feart.2022.927059
Bosio, G. & Bianucci, G. & Collareta, A. & Landini, W. & Urbina, M. & Di Celma, C. (2022) Ultrastructure, composition, and Sr-87/Sr-86 dating of shark teeth from lower Miocene sediments of southwestern Peru. Journal of South American Earth Sciences, 118, Article 103909 https://dx.doi.org/10.1016/j.jsames.2022.103909
Ferron, H.G. & Ballell, A. & Botella, H. & Martinez-Perez, C. (2022) Biomechanics of Machaeracanthus pectoral fin spines provide evidence for distinctive spine function and lifestyle among early chondrichthyans. Journal of Vertebrate Paleontology, 41(6), Article e2090260 https://dx.doi.org/10.1080/02724634.2021.2090260
Hovestadt, D.C. (2022)  A partial skeleton of Carcharias cuspidatus including embryos from the Oligocene of Germany. Cainozoic Research, 22(1), 25–36
Janvier, P. (2022) Description des restes d'Elasmobranches (Pisces) du Dévonien moyen de Bolivie. Palaeovertebrata, 7(4), 126–132
Kim, S.L. & Yeakel, J.D. & Balk, M.A. & Eberle, J.J. & Zeichner, S. & Fieman, D. & Kriwet, J. (2022) Decoding the dynamics of dental distributions: insights from shark demography and dispersal. Proceedings of the Royal Society of London, Series B, 289(1977), Article 20220808 https://dx.doi.org/10.1098/rspb.2022.0808
Kuhn, E. (2022) Fischfunde aus dem Jura der Umgebung von Baden (Aargau). Eclogae Geologicae Helvetiae, 34, 297–302
Mottequin, B. & Fischer, J. & Goolaerts, S. & Olive, S. (2022) Revisiting the chondrichthyan egg capsules inventory from the Pennsylvanian (Carboniferous) of Belgium: new data and perspectives. The Science of Nature, 109(4), Article 39  https://dx.doi.org/10.1007/s00114-022-01809-6
Rudebeck, B. & Riehmann, H. (2022) Fossile hajtaender fra Trelde Naes. Self-published
Zazzera, A. & Girone, A. & La Perna, R. & Marino, M. & Maiorano, P. & Sardella, R. & Montenegro, V. & Francescangeli, R. & Bianucci, G. (2022) Systematics, taphonomy and palaeobiogeography of a balaenopterid (Cetacea, Mysticeti) from the Early Pleistocene of southern Italy. Geobios, 71, 51–65 https://dx.doi.org/10.1016/j.geobios.2022.01.001

Beveridge, I. & Bennett, M.B. (2022) The genus Hemionchos Campbell & Beveridge, 2006 (Cestoda: Trypanorhyncha) in Australian manta rays (Batoidea: Mobulidae). Transactions of the Royal Society of South Australia, in press https://dx.doi.org/10.1080/03721426.2022.2096196
Boxshall, G.A. & Barton, D.P. & Kirke, A. & Zhu, X.C. & Johnson, G. (2022) Two new parasitic copepods of the family Sphyriidae (Copepoda: Siphonostomatoida) from Australian elasmobranchs. Systematic Parasitology, in press https://dx.doi.org/10.1007/s11230-022-10054-4
Franzese, S. & Mutti, L.D. & Tropea, C. & Ivanov, V.A. (2022) Morphological study of members of the genus Echeneibothrium (Cestoda: Rhinebothriidea: Echeneibothriidae) from rajiform skates of the Argentine Sea and analysis of the phylogenetic relationships within the family Echeneibothriidae. Zoologischer Anzeiger, 299, 1–10 https://dx.doi.org/10.1016/j.jcz.2022.05.002



Associate Professor Nicolas Straube at the Department of Natural History, University Museum shows the jaw of the shark species Somniosus microcephalus. Photo/illus.: Siri Skretting Jansen, UM


Hvithaier kan ha bidratt til utryddelse av den aller største haien som har levd!

Ny metode for undersøkelse av sink-isotoper i tenner fra den utdødde giganthaien Otodus megalodon og tenner fra fossile og moderne hvithai gir forskerne helt ny kunnskap.


New research shows 2/3 of species in global shark fin trade at risk of extinction

Researchers say coastal sharks could benefit from increased protections, international trade regulations

Date: July 20, 2022
Source: Mote Marine Laboratory
Summary: Researchers sampled nearly 10,000 shark fin trimmings from markets in Hong Kong -- one of the largest shark fin trade hubs in the world. With a little DNA detective work, they unraveled the mystery of what fin belonged to what species, and found that endangered or threatened species were disproportionately represented in the trade, highlighting the need for additional international regulations in trade and shark management practices.

How 'shark week' could inspire love for ocean predators

Date: July 21, 2022
Source: North Carolina State University
Summary: Findings from a new study indicate more positive YouTube videos could be a useful tool for shark conservation.


New IUCN Shark News Newsletter is out!
Download: https://www.iucnssg.org/shark-news.html