NEWSLETTER 06+07/2021 15.07.2021

Please acknowledge use of the database www.shark-references.com in your publications, and cite: 

Pollerspöck, J. & Straube, N. 2021, Bibliography database of living/fossil sharks, rays and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali), www.shark-references.com, World Wide Web electronic publication, Version 2021


STRAUBE, N. & LYRA, M.L. & PAIJMANS, J.L.A. & PREICK, M. & BASLER, N. & PENNER, J. & RÖDEL, M.-O. & WESTBURY, M.V. & HADDAD, C.F.B. & BARLOW, A. & HOFREITER, M. (2021): Successful application of ancient DNA extraction and library construction protocols to museum wet collection specimens. Molecular Ecology Resources, in press

Abstract: Millions of scientific specimens are housed in museum collections, a large part of which are fluid preserved. The use of formaldehyde as fixative and subsequent storage in ethanol is especially common in ichthyology and herpetology. This type of preservation damages DNA and reduces the chance of successful retrieval of genetic data. We applied ancient DNA extraction and single stranded library construction protocols to a variety of vertebrate samples obtained from wet collections and of different ages. Our results show that almost all samples tested yielded endogenous DNA. Archival DNA extraction was successful across different tissue types as well as using small amounts of tissue. Conversion of archival DNA fragments into single-stranded libraries resulted in usable data even for samples with initially undetectable DNA amounts. Subsequent target capture approaches for mitochondrial DNA using homemade baits on a subset of 30 samples resulted in almost complete mitochondrial genome sequences in several instances. Thus, application of ancient DNA methodology makes wet collection specimens, including type material as well as rare, old or extinct species, accessible for genetic and genomic analyses. Our results, accompanied by detailed step-by-step protocols, are a large step forward to open the DNA archive of museum wet collections for scientific studies.

ADNET, S. & FEICHTINGER, I. & HARZHAUSER, M. & POLLERSPÖCK, J. (2021) A mesopelagic selachian fauna from the middle Eocene of St. Pankraz (Austria) reveals homogeneity in deep-marine environments during the warm period in Europe. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, 301 (1): 25-63    

Abstract: Repeated bulk sampling for over a decade in an indurated glauconitic sandy marl horizon at St. Pankraz Salzburg, Austria, has yielded a diverse assemblage of 37 elasmobranchs (sharks and rays) from the early middle Eocene (Lutetian). This elasmobranch fauna is dominated by epipelagic and mesopelagic taxa known today to preferentially inhabit the middle or outer continental shelf and upper slope, indicating that the depositional environment of the top Member of Kressenberg Formation in Austria has a more complex bathymetric history than previously thought. As these new occurrences fill a substantial gap in the sporadic fossil record of Eocene mesopelagic elasmobranchs, comparisons of this assemblage with the coeval mesopelagic faunas indicate that this northwestern Tethyan realm association shares considerable similarities with those recovered from the North Sea Basin and the northeastern Atlantic. This suggests that the faunal homogeneity observed in neritic and coastal elasmobranch communities during the warm early middle Eocene is also characterised in mesopelagic habitats.


New database report published by team shark-references!
- You want to know how many species of sharks, rays and chimaeras there are?
- You want to know the distribution of species among the orders or families?
- You want to know the references of the first descriptions?
- You want a list of all sharks, rays and chimaeras ever described with their synonyms?

Our new data report can answer all these questions for you!
Abstract: The table and provided download links below are intended for informational use in Chondrichthyan research. The allocation aims for faciliating to find species numbers and most recent information on taxonomic changes. We will regularly update the table and download links at lest twice annually. The updates will be announced on facebook (https://www.facebook.com/sharkreferences) and in our monthly newsletter (sign up here: https://eepurl.com/sJNGb). The Excel sheet allows for the application of individual filter- and sorting options. The list of described spsecies complements taxonomic information for the list of valid species by providing synonyms and / or new taxonomic combinations.


Our Co-Editor Nico was recently invited to join the the IUCN shark specialist group. We are very happy about that and will support the IUCN and their work the best we can!
Check out:
Web: https://www.iucnssg.org/
FB: https://www.facebook.com/IUCNShark


Maria Cristina Oddone, Instituto de Ciências Biológicas, Universidade Federal de Rio Grande, Brasil sent me this two obituaries written in memory of professor Carolus Maria Vooren.

Obituary: Professor Carolus Maria Vooren (1941-2021).
Pan-American Journal of Aquatic Sciences, 16(1): 1-4.

OBITUÁRIO - CAROLUS MARIA VOOREN (14/11/1941-12/3/2021)
v. 54 n. 1 (2021): Arquivos de Ciências do Mar

Would you like to become a shark-reference partner? Please contact us per E-mail!

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New Images

Many thanks to the following people for providing images:

Frederik Mollen (Elasmobranch Research Belgium) for the images of Carcharhinus albimarginatus

Spencer Lucas for some images of Dracopristis hoffmanorum HODNETT, GROGAN, LUND, LUCAS, SUAZO, ELLIOTT & PRUITT, 2021

Dr. Simon Weigmann, Elasmobranch Research Laboratory, Hamburg, Germany for a images of Etmopterus broseiAcroteriobatus andysabini and Acroteriobatus stehmanni

Bryan Huerta for a image of Carcharhinus brevipinnaCarcharhinus isodon and Sphyrna tiburo

Adam Anderson for pics of Notorynchus primigeniusChlamydoselachus keyesiGladioserratus aptiensis and  Notidanodon lanceolatus

LOBODA, LASSO, ROSA & DE CARVALHO for an images of Paratrygon orinocensis and Paratrygon parvaspina

Iris Feichtinger and Viola Winkler, Wien for images of Cladodus gailensis

Gilles Cuny for a image of Triodus aeduorum

Md Jayedul Islam, Dhaka, Bangladesh for a image of Glaucostegus younholeei

Bianca S. Rangel, Institute of Biosciences, University of São Paulo for images of Pseudobatos horkelii

Missing papers:

Many thanks to all friends of shark-references, who sent us some missing papers last month!

Shark-References would kindly like to ask you for your contribution to this project.

At the moment we are looking for some of the following papers:

Extinct Chondrichthyes:

MATSUMOTO, H. (1936) Upper Miocene vertebrates from Kumanodô, Natori district, province of Rikuzen. Dobutsugaku Zasshi, 48: 475–480, 5 fig.

ALVINERIE, J. & ANDREIEFF, P. & ANGLADA, R. & AUBERT, J. & CAPPETTA, H. & CARALP, M. & CARATINI, C. & CARBONNEL, G. & CATZIGRAS, F. & COURME-RAULT, M.-D. & CHATEAUNEUF, J.-J. & DEMARCQ, G. & DUCASSE, O. & FATTON, E. & GLAÇON, G. & LABRACHERIE, M. & LAURIAT, A. & LE CALVEZ, Y. & LORENZ, C. & MAGNE, J. & MARGEREL, J.-P. & POIGNANT, A. & PUJOL, C. & ROGER, J. & ROMAN, J. & BLONDEAU, A. & MULLER, C. (1973) A propos de la limite oligo-miocène: résultats préliminaires d'une recherche collective sur les gisements d'Escornébéou (Saint-Géours-de-Maremne, Landes, Aquitaine méridionale). Présence de Globigerinoides dans les faunes de l'Oligocène supérieur. Comptes rendus sommaires des séances de la Société géologique de France: 75–76

Extant Chondrichthyes:

KAMOHARA, T. (1943) Some unrecorded and two new fishes from Prov. Tosa, Japan. Bulletin of the Biogeographical Society of Japan, 13 (17): 125–137

DE BUEN, F. (1950) Contribuciones a la Ictiología. II. El tiburón vitamínico de la costa uruguaya Galeorhinus vitaminicus nov. sp., y algunas consideraciones generales sobre su biología. Publicaciones Cientificas, Servicio Oceanografico y de Pesca, Ministerio de Industrias y Trabajo, Montevideo No. 4: 153–162.

WEIBEZAHN, F.H. (1953) Una nueva especie de Scyliorhinus de Venezuela (Chondrichthyes - Elasmobranchii). Novedades cientificas. Serie zoológica. Museo de Historia Natural La Salle, 9: 1–7.

SMITH, J.L.B. (1958) The mystery killer, the new shark Carcharhinus vanrooyeni. Veld & Vlei, 3 (9): 12–14, 28.

GUBANOV, E.P. & SCHLEIB, N.A. (1980) Sharks of the Arabian Gulf. Kuwait Ministry of Public Works, Agracultural Department, Fisheries Division. Sharks of the Arabian Gulf.: 1–69

DOLGANOV, V.N. (1983) Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod. [Manual for identification of cartilaginous fishes of Far East seas of USSR and adjacent waters.] TINRO, Vladivostok. Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod.: 92 pp.

Please support www.shark-references.com and send missing papers (not listed papers or papers without the info-symbol) to juergen.pollerspoeck@shark-references.com or nicolas.straube@shark-references.com



Upcoming Meetings:


EEA Meeting 2021 in Leiden, Netherlands

Dear friends and colleagues,
We are very pleased to inform you that abstract submission for EEA2021 is available online at https://www.elasmobranch.nl/eea2021/abstract-submission/ 
Registration will follow in due course, so keep an eye on the website: https://www.elasmobranch.nl/eea2021/ 

The 24th Annual Scientific Meeting of the European Elasmobranch Association is planned to be held by the Dutch member group Nederlandse Elasmobranchen Vereniging (NEV) in Leiden, the Netherlands between 3rd and 5th November, 2021.

NEV, in consultation with the EEA Board, has decided to tentatively organise this year’s EEA Meeting as an in-person meeting. Luckily, NEV was able of having secured the same prime venue at the fascinating Naturalis Biodiversity Center! Leiden is a beautiful old university town just 20 mins from Schiphol airport and international train station.

Prior to the main meeting we will organise an online session on November 2nd to allow those who cannot attend in person to contribute.

We understand that there is still a lot of uncertainty as far as travel etc. is concerned. In mid-July we will make the decision whether or not to host a live event.

If an in-person meeting is not possible the entire EEA2021 will be online.

In the coming weeks the website for early registration will be updated, as well as opening the abstract submission for either an online or live presentation.

Details can be found on the NEV website: https://www.elasmobranch.nl/eea2021/


Welcome to ESEB 2021 Congress.

Given the current situation, the organizers
decided to postpone the congress by a year.

Welcome to the ESEB 2021 congress, to be held at the Prague Congress Centre, the Czech Republic, on 22-27 August 2021.

The congresses of the European Society for Evolutionary Biology (ESEB) are organized biennially since 1987 and are now among the largest conferences in evolutionary biology with about 1400 – 1700 participants.

The congress will commence on Sunday, August 22, 2021, with the welcome reception, and will continue until Friday, August 27, concluding with the conference dinner at Občanská Plovárna Restaurant on this evening.

The Prague Congress Centre is easily accessible from the city centre and has a beautiful panoramic view of Prague.  



92nd Annual Meeting of the Paläontologische Gesellschaft!

This year's Annual Meeting will be held as online conference from September 27 to 30, 2021.

We are pleased to host the 92nd annual conference of the Paläontologische Gesellschaft (PalGes). The meeting, like the previous ones, is designed as an international meeting to present cutting-edge research from palaeobiology, palaeontology, geobiology and related subjects. The Paläontologische Gesellschaft is one of the oldest and largest palaeontological societies in the world and we "Viennese" are now hosting the annual meeting for the fifth time after 1923, 1954, 1963 and 2011.

The venue will be held as online conference only due to uncertainties over travel and meeting size restrictions related to COVID-19 development. Participation in the conference is free of charge. Unfortunately, the online format means that there will not be any poster session or conference dinner this year.

We will keep you up to date with new and exciting details of our virtual meeting. If you have questions, please do not hesitate to contact us via e-mail.





Extant Chondrichthyes:

EBERT, D.A. & LESLIE, R.W. & WEIGMANN, S. (2021):
 Etmopterus brosei sp. nov.: a new lanternshark (Squaliformes: Etmopteridae) from the southeastern Atlantic and southwestern Indian oceans, with a revised key to the Etmopterus lucifer clade. Marine Biodiversity, 51: 53
New species: Etmopterus brosei
Abstract: A new species of lanternshark, Etmopterus brosei sp. nov. (Squaliformes: Etmopteridae), is described from the southeastern Atlantic and southwestern Indian oceans. The new species resembles other members of the Etmopterus lucifer Jordan & Snyder, 1902 clade in having linear rows of dermal denticles, and most closely resembles the conspecific E. sculptus Ebert, Compagno, & De Vries, 2011 from the southeastern Atlantic and southwestern Indian oceans. The new species is fairly common along the upper continental slopes off South Africa, Mozambique, and seamounts along the Madagascar Ridge, including Walters Shoal, in 480–1200 m depth. It can be distinguished from other members of the E. lucifer clade by a combination of characteristics, including the arrangement of flank and caudal markings, shape and size of flank marking, the arrangement of dermal denticles along the body, and the presence of dermal denticles on the dorsal fin bases. A revised key to the Etmopterus lucifer clade is provided.
WHITE, W.T. & FRICKE, R. (2021): Raja mauritaniensis: a replacement name for Raja africana Capapé, 1977 (Rajiformes: Rajidae), a junior homonym of Raja africana Bloch & Schneider, 1801 (Myliobatiformes: Dasyatidae). Zootaxa, 4970 (2): 399–400
New name: Raja mauritaniensis
AbstractRaja africana Capapé, 1977 is a primary junior synonym of Raja africana Bloch & Schneider, 1801 and therefore permanently invalid (International Code of Zoological Nomenclature, article 57.2) and must be replaced. Raja africana Bloch & Schneider, 1801 was first described by Bloch & Schneider (1801: 367), based on a specimen from Guinea, West Africa (eastern Atlantic Ocean). The unique holotype is extant in the Zoologisches Museum of the Humboldt University, Berlin (ZMB 7837, a partial dry skin). The species was treated as valid as Urogymnus africanus (Bloch & Schneider 1801) by Compagno & Roberts (1984: 285), but later synonymized with Urogymnus asperrimus (Bloch & Schneider 1801) in the subfamily Urogymninae of the family Dasyatidae (Myliobatiformes) by Compagno (1986: 141), Capapé & Desoutter (1990: 63) and Séret (2016: 1418). It is widespread in the eastern Atlantic, Red Sea and Indo–West Pacific.

LOBODA, T.S. & LASSO, C.A. & ROSA, R.S & DE CARVALHO, M.R. (2021): Two new species of freshwater stingrays of the genus Paratrygon (Chondrichthyes: Potamotrygonidae) from the Orinoco basin, with comments on the taxonomy of Paratrygon aiereba. Neotropical Ichthyology, 19 (2): e200083
New species: Paratrygon orinocensisParatrygon parvaspina
Abstract: The genus Paratrygon, currently recognized as the sole monotypic genus of the family Potamotrygonidae, has a considerably greater diversity than previously indicated, including molecular studies, which supported P. aiereba (hitherto the only recognized species in the genus) as a possible species complex. Here we describe two new species of the genus that are both endemic to and sympatric in the Orinoco basin. Paratrygon aiereba, type species of the genus, is now restricted to the Amazon basin. Both new species are identified and defined through morphological characters such as coloration, dermal denticle morphology, arrangement of thorns, distribution and morphology of ventral lateral line canals, morphology of skeletal elements, and morphometrics. An extensive comparison of these characters between the new species herein described and P. aiereba is presented. Finally, a taxonomic reappraisal of P. aiereba is provided through a revision of preserved material and its original description, plus new evidence about its type-locatity, collectors, and a reconsideration of the destination of its type-specimen.

WEIGMANN, S. & EBERT, D.A. & SÉRET, B. (2021): Resolution of the Acroteriobatus leucospilus species complex, with a redescription of A. leucospilus (Norman, 1926) and descriptions of two new western Indian Ocean species of Acroteriobatus (Rhinopristiformes, Rhinobatidae). Marine Biodiversity, 51: 58
New species: Acroteriobatus andysabini, Acroteriobatus stehmanni
Abstract: Recent sampling efforts and examinations of museum material provided evidence for a complex of species within Acroteriobatus leucospilus (Norman, 1926). The present manuscript contains a redescription of A. leucospilus involving the syntypes and additional material, as well as formal descriptions of two new species of Acroteriobatus Giltay, 1928. All specimens of both new species were found in the western Indian Ocean. Individuals of the first new species, hereafter referred to as Acroteriobatus andysabini sp. nov., were identified originating from Madagascar, and specimens of the second new species, hereafter referred to as Acroteriobatus stehmanni sp. nov., were only found off Socotra Islands at the junction between the Gulf of Aden and the Arabian Sea. Both new species appear to be endemic to the respective region and allopatric to A. leucospilus and occur in coastal waters to at least 80 m and 43 m depth, respectively. They differ from A. leucospilus in a number of characteristics including the maximum size and coloration in fresh. Acroteriobatus andysabini sp. nov. is a large species growing to more than 100 cm total length (TL) and with elongated bluish-gray spots on snout giving “stripe-nosed” appearance; numerous small bluish-gray spots on pectoral, pelvic, dorsal, and caudal fins; brown spots on trunk and fin bases; lateral tail folds striped orange and white; and ventral surface largely white but with a V-shape pattern of faint to dark speckled black spots on snout tip. Acroteriobatus stehmanni sp. nov. is a small species growing to ~62 cm TL and with sparse patterning with small bluish-gray circular spots confined to snout tip, posterior pectoral-fin margins, a pair on midbody, and few on posterior pelvic-fin margins, rather indistinct small to larger dark brown spots, and lateral tail folds and ventral surface white. Acroteriobatus leucospilus is a medium-sized species growing to ~96 cm TL and with patterning similar to A. andysabini sp. nov. but ventral surface uniformly white and lateral tail folds white or striped blue and brown. Taxonomical differences include nasal lamellae counts (42–48 in A. andysabini sp. nov. vs. 43–48 in A. stehmanni sp. nov. vs. 37–41 in A. leucospilus), snout angle (76–85° vs. 71–77° vs. 68–81°), and dorsal head length (24.2–33.5% vs. 17.2–22.8% TL vs. 24.0–29.2% TL). A key to the species of Acroteriobatus is given for the first time.

HABIB, K.A. & ISLAM, M.J. (2021): Description of a new species of giant guitarfish, Glaucostegus younholeei sp. nov. (Rhinopristiformes: Glaucostegidae) from the northern Bay of Bengal, Bangladesh. Zootaxa, 4995 (1): 129–146
New speciesGlaucostegus younholeei
Abstract: A new species of giant guitarfish, Glaucostegus younholeei sp. nov., is described from 13 specimens, 730–933 mm total length, collected from fish landing center of Bangladesh Fisheries Development Corporation in Cox’s Bazar district of Bangladesh. The new species is distinguished from congeners in having the following combination of characters: Body brownish or greyish in color with a narrowly wedge-shaped disc, and long narrow bluntly pointed snout (angle 31–40°), and broad oblique nostrils with the narrow anterior opening. Nostrils about half of the mouth width, subequal (0.98–1.33) to internasal width; ~55–57 nasal lamellae; anterior nasal flaps slightly penetrating into internasal space, their interspace 2.20– 2.61 in length of the posterior nasal aperture. Orbit very small in adults, diameter 8.19–11.62 in preorbital length, 2.25–2.69 in interorbital space. Rostral ridges almost joined along their entire length; margin of cranium sharply demarcated before eyes. Spiracular folds very short and widely separated. Skin rough, densely covered with small denticles, more coarsely granular on the dorsal surface than ventrally, enlarged between orbits and in a distinct band between nape and first dorsal fin. Tail relatively longer, length 1.15–1.48 in disc length; dorsal fins narrowly spaced, interspace 1.32–2.11 in base length of the first dorsal fin. Clasper length in adult male 4.37–5.70 in total length. Phylogenetic analysis of DNA barcode sequences also shows the clear divergence of Glaucostegus younholeei from other congeneric species obtained from GenBank. A key is provided to the 8 known members including new species of the genus Glaucostegus.

Extinct Chondrichthyes:

STUMPF, S. & ETCHES, S. & UNDERWOOD, C.J. & KRIWET, J. (2021): Durnonovariaodus maiseyi gen. et sp. nov., a new hybodontiform shark-like chondrichthyan from the Upper Jurassic Kimmeridge Clay Formation of England. Peerj, 9: e11362
New genus: Durnonovariaodus
New species: Durnonovariaodus maiseyi
Abstract: A partial skeleton of a hybodontiform shark-like chondrichthyan from the Upper Jurassic Kimmeridge Clay Formation of Dorset, England, is described and designated as a new genus and species, Durnonovariaodus maiseyi gen. et sp. nov. The holotype and only known specimen, which is represented by disarticulated splanchnocranial elements with associated teeth, a single dorsal fin spine, the pelvic girdle, as well as unidentifiable cartilage fragments, plus countless dermal denticles, exhibits a puzzling combination of dental and skeletal features, providing important new insights into the morphological and ecological diversity of hybodontiforms. Durnonovariaodus gen. nov. displays a unique set of dental characters, showing close morphological resemblance to Secarodus from the Middle Jurassic of England, which was erected for distinctive, strongly labio-lingually compressed multicuspid cutting teeth originally described as Hybodus polyprion. Skeletally, Durnonovariaodus gen. nov. resembles Hybodus and Egertonodus in having a palatoquadrate with a palatobasal process and an ethmoidal articular surface, combined with the possession of dorsal fin spines ornamented with costae. Therefore, and given the absence of any conclusive phylogenetic framework, Durnonovariaodus maiseyi gen. et sp. nov. is here tentatively referred to Hybodontidae until more complete material becomes available in order to enable a more reliable suprageneric identification. The holotype of Durnonovariaodus maiseyi gen. et sp. nov. contains two separate pelvic half-girdles, a feature previously considered as evolutionarily primitive among hybodontiforms. However, unfused pelvic half-girdles also occur in the supposedly closely related species Hybodus hauffianus and may in fact have been more widely distributed among hybodontiforms than previously thought, thus rendering the phylogenetic utility of separated pelvic half-girdles for inferring hybodontiform interrelationships difficult and unresolved.

FEICHTINGER, I. & IVANOV, A.O. & WINKLER, V. & DOJEN, C. & KINDLIMANN, R. & KRIWET, J. & PFAFF, C. & SCHRAUT, G. & STUMPF, S. (2021): Scarce ctenacanthiform sharks from the Mississippian of Austria with an analysis of Carboniferous elasmobranch diversity in response to climatic and environmental changes. Journal of Vertebrate Paleontology, in press
New species: Cladodus gailensis
Abstract: The Carboniferous is characterized by drastic climatic and environmental fluctuations, which include multiple phases of glaciation resulting in an icehouse climate. Additionally, dynamic continental reconfigurations forced the contraction of the Rheic Ocean resulting in the closure of the Rheic–Tethyan Gateway, which precluded further faunal exchanges between the North American and Eurasian marine realms. Interestingly, cartilaginous fishes seem to be relatively immune to these drastic climatic and environmental changes. The Eurasian fossil record of Paleozoic sharks is strongly biased towards intensively sampled localities from England, Ireland, Scotland, and the Russian Platform. Here we present rare dental material from the Serpukhovian (early Carboniferous) of Austria, adding new information to the paleogeographic distribution of ctenacanthiform sharks. The new material revealed the first record of the genus Saivodus in Central Europe and allowed us to recognize a new species, Cladodus gailensis sp. nov., and a remnant of fossilized cartilage. In an attempt to identify possible linkages between climatic or environmental fluctuations on shark diversity throughout the Carboniferous, we provide a synopsis of the distribution and diversity of elasmobranchs based on primary literature. This preliminary assessment at genus level indicates two pronounced events of extinction, with the first one occurring during the latest Mississippian and the second one towards the end of the Pennsylvanian. The first extinction event distinctly correlates with the known diversity decline of other marine inhabitants and the second occurred during an unstable period of multiple phases of glaciation.

LUCCISANO, V. & PRADEL, A. & AMIOT, R. & GAND, G. & STEYER, J.-S. & CUNY, G. (2021): A new Triodus shark species (Xenacanthidae, Xenacanthiformes) from the lowermost Permian of France and its paleobiogeographic implications. Journal of Vertebrate Paleontology, in press
New species: Triodus aeduorum
Abstract: The Xenacanthiformes from Central and Eastern European deposits have been extensively studied, but the systematics of the species from the French Carboniferous–Permian Autun Basin (Saône-et-Loire) remains debated. Numerous xenacanthiform remains are still identified under the doubtful genus ‘Expleuracanthus’, and many of them consist of isolated dorsal spines which are difficult to identify. Numerous well-preserved specimens are still undescribed and the diversity of the xenacanthiform fauna from the Autun Basin is poorly understood. For example, specimens of the genus Triodus from the Muse oil-shale bed (OSB) of the Autun Basin have no specific attribution, whereas this genus is widely distributed across European Carboniferous–Permian basins. In this study, we describe new specimens of Triodus from the lowermost Permian of the Muse OSB. They allow the erection of a new species, Triodus aeduorum sp. nov., and discussion of the validity of several species from the same locality: ‘Expleuracanthus’ frossardi is considered as a nomen dubium and other Triodus specimens need for the time being to be left in open nomenclature as Triodus sp. These results highlight the endemism of the Triodus species in each European Carboniferous–Permian basin and raise the question of how they migrated from one to another.


no taxonomic news this month


PLEASE send your new papers to
juergen.pollerspoeck@shark-references.com or 

Latest Research Articles

Extant Chondrichthyes:
AHMED, M.S. & DATTA, S.K. & SAHA, T. & HOSSAIN, Z. (2021) Molecular characterization of marine and coastal fishes of Bangladesh through DNA barcodes. Ecology and Evolution, 11 (9): 3696-3709 https://dx.doi.org/10.1002/ece3.7355
ARASTEHFAR, S. & CHEW, C.M. (2021) Effects of root chord movement on thrust generation of oscillatory pectoral fins. Bioinspiration & Biomimetics, 16 (3): 036009 https://dx.doi.org/10.1088/1748-3190/abc86b
BAKIU, R. & SOLDO, A. (2021) Shark capture by commercial fisheries in Albania. Journal of Applied Ichthyology, in press https://dx.doi.org/10.1111/jai.14208
BASS, N.C. & DAY, J. & GUTTRIDGE, T.L. & KNOTT, N.A. & BROWN, C. (2021) Intraspecific variation in diel patterns of rocky reef use suggests temporal partitioning in Port Jackson sharks. Marine and Freshwater Research, in press https://dx.doi.org/10.1071/mf20204
BERIO, F. & BROYON, M. & ENAULT, S. & PIROT, N. & LOPEZ-ROMERO, F.A. & DEBIAIS-THIBAUD, M. (2021) Diversity and Evolution of Mineralized Skeletal Tissues in Chondrichthyans. Frontiers in Ecology and Evolution, 9: 660767 https://dx.doi.org/10.3389/fevo.2021.660767
BERNAL-PÉREZ, S. & OCHOA-BÁEZ, R.I. & GALVÁN-MAGAÑA, F. & SOTO-LÓPEZ, K. (2021) Reproductive biology of the Swell Shark Cephaloscyllium ventriosum (Carcharhiniformes: Scyliorhinidae). Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14765
BESNARD, L. & LE CROIZIER, G. & GALVÁN-MAGAÑA, F. & POINT, D. & KRAFFE, E. & KETCHUM, J. & MARTINEZ RINCON, R.O. & SCHAAL, G. (2021) Foraging depth depicts resource partitioning and contamination level in a pelagic shark assemblage: Insights from mercury stable isotopes. Environmental Pollution, 283: 117066 https://dx.doi.org/10.1016/j.envpol.2021.117066
BIRKMANIS, C.A. & SIMMONS, L.W. & SEQUEIRA, A.M.M. (2021) Response to Limitations on inferring shark vulnerability from spatial habitat protection. Global Ecology and Conservation, 26: e01466 https://dx.doi.org/10.1016/j.gecco.2021.e01466
BONFIL, R. & PALACIOS-BARRETO, P. & VARGAS, O.U.M. & RICANO-SORIANO, M. & DIAZ-JAIMES, P. (2021) Detection of critically endangered marine species with dwindling populations in the wild using eDNA gives hope for sawfishes. Marine Biology, 168 (5): 60 https://dx.doi.org/10.1007/s00227-021-03862-7
BOUVEROUX, T. & LOISEAU, N. & BARNETT, A. & MAROSI, N.D. & BRUNNSCHWEILER, J.M. (2021) Companions and Casual Acquaintances: The Nature of Associations Among Bull Sharks at a Shark Feeding Site in Fiji. Frontiers in Marine Science, 8: 678074 https://dx.doi.org/10.3389/fmars.2021.678074
BOUYOUCOS, I.A. & TRUJILLO, J.E. & WEIDELI, O.C. & NAKAMURA, N. & MOURIER, J. & PLANES, S. & SIMPFENDORFER, CA. & RUMMER, J.L. (2021) Investigating links between thermal tolerance and oxygen supply capacity in shark neonates from a hyperoxic tropical environment. Science of the Total Environment, 782: 146854 https://dx.doi.org/10.1016/j.scitotenv.2021.146854
BOWLBY, H.D. & BENOIT, H.P. & JOYCE, W. & SULIKOWSKI, J. & COELHO, R. & DOMINGO, A. & CORTES, E. & HAZIN, F. & MACIAS, D. & BIAIS, G. & SANTOS, C. & ANDERSON, B. (2021) Beyond Post-release Mortality: Inferences on Recovery Periods and Natural Mortality From Electronic Tagging Data for Discarded Lamnid Sharks. Frontiers in Marine Science, 8: 619190 https://dx.doi.org/10.3389/fmars.2021.619190
BRACCINI, M. & HESP, A. & MOLONY, B. (2021) Risk-based weight of evidence assessment of commercial sharks in western Australia. Ocean & Coastal Management, 205: 105501 https://dx.doi.org/10.1016/j.ocecoaman.2020.105501
BRADSHAW, C.J.A. & MEAGHER, P. & THIELE, M.J. & HARCOURT, R.G. & HUVENEERS, C. (2021) Predicting potential future reduction in shark bites on people. Royal Society Open Science, 8 (3): 201197 https://dx.doi.org/10.1098/rsos.201197
BUDDLE, A.L. & VAN DYKE, J.U. & THOMPSON, M.B. & SIMPFENDORFER, C.A. & MURPHY, C.R. & DOWLAND, S.N. & WHITTINGTON, C.M. (2021) Structure of the paraplacenta and the yolk sac placenta of the viviparous Australian sharpnose shark, Rhizoprionodon taylori. Placenta, 108: 11-22 https://dx.doi.org/10.1016/j.placenta.2021.03.010
CAMBRA, M. & MADRIGAL-MORA, S. & CHINCHILLA, I. & GOLFÍN-DUARTE, G. & LOWE, C.G. & ESPINOZA, M. (2021) First record of a potential neonate tiger shark (Galeocerdo cuvier) at a remote oceanic island in the Eastern Tropical Pacific. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14774
CAMPBELL, M.J. & MCLENNAN, M.F. & COURTNEY, A.J. & SIMPFENDORFER, C.A. (2021) Life-history characteristics of the eastern shovelnose ray, Aptychotrema rostrata (Shaw, 1794), from southern Queensland, Australia. Marine and Freshwater Research, in press https://dx.doi.org/10.1071/mf20347
CANEDO-APOLAYA, R.M. & ORTIZ-ALVAREZ, C. & ALFARO-CORDOVA, E. & ALFARO-SHIGUETO, J. & VELEZ-ZUAZO, X. & MANGEL, J.C. & SICCHA-RAMIREZ, R. & YAMASHIRO, C. & RAMIREZ, J.L. (2021) Species Delimitation of Southeast Pacific Angel Sharks (Squatina spp.) Reveals Hidden Diversity through DNA Barcoding. Diversity-Basel, 13 (5): 177 https://dx.doi.org/10.3390/d13050177
CASTELLANOS-GALINDO, G.A. & HERRON, P. & NAVIA, A.F. & BOOTH, H. (2021) Shark conservation and blanket bans in the eastern Pacific Ocean. Conservation Science and Practice, in press https://dx.doi.org/10.1111/csp2.428
CASTILLO, C.G.A & MORALES, O.S. (2021) Characterization of the artisanal elasmobranch fisheries off the Pacific coast of Guatemala. Fishery Bulletin, 119 (1): 3-9 https://dx.doi.org/10.7755/FB.119.1.2
CHAN, M.Y.C. &SOSA-NISHIZAKI, O. & PÉREZ-JIMÉNEZ, J.C. (2021) Potential distribution of critically endangered hammerhead sharks and overlap with the small-scale fishing fleet in the southern Gulf of Mexico. Regional Studies in Marine Science, 46: 101900 https://dx.doi.org/10.1016/j.rsma.2021.101900
CHAPMAN, D.D. & ALI, K. & MACNEIL, M.A. & HEUPEL, M.R. & MEEKAN, M. & HARVEY, E.S. & SIMPFENDORFER, C.A. & HEITHAUS, M.R. (2021) Long-term investment in shark sanctuaries. Science, 372 (6541): 473 https://dx.doi.org/10.1126/science.abj0147
COLBACHINI, H. & PIZZUTTO, C.S. & MESQUITA, L.M.D. & GADIG, O.B.F. (2021) Environmental enrichment effects on the reproductive behavior of captive nurse sharks Ginglymostoma cirratum. Environmental Biology of Fishes, in press https://dx.doi.org/10.1007/s10641-021-01087-7
COLEMAN, N.C. & BURGE, E.J. (2021) Association behavior between sand tiger sharks and round scad is driven by mesopredators. Peerj, 9: e11164 https://dx.doi.org/10.7717/peerj.11164
COLLINS, C. & NUNO, A. & BRODERICK, A. & CURNICK, D.J. & DE VOS, A. & FRANKLIN, T. & JACOBY, D.M.P. & MEES, C. & MOIR-CLARK, J. & PEARCE, J. & LETESSIER, T.B. (2021) Understanding Persistent Non-compliance in a Remote, Large-Scale Marine Protected Area. Frontiers in Marine Science, 8: 650276 https://dx.doi.org/10.3389/fmars.2021.650276
CONTRERAS-REYES, J.E. & WIFF, R. & SOTO, J. & DONOVAN, C.R. & ARAYA, M. (2021) Biphasic growth modelling in elasmobranchs based on asymmetric and heavy-tailed errors. Environmental Biology of Fishes, 104 (5): 615-628 https://dx.doi.org/10.1007/s10641-021-01100-z
COOPER, M.K. & HUERLIMANN, R. & EDMUNDS, R.C. & BUDD, A.M. & LE PORT, A. & KYNE, P.M. & JERRY, D.R. & SIMPFENDORFER, C.A. (2021) Improved detection sensitivity using an optimal eDNA preservation and extraction workflow and its application to threatened sawfishes. Aquatic Conservation: Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/aqc.3591
CROBE, V. & FERRARI, A. & HANNER, R. & LESLIE, R.W. & STEINKE, D. & TINTI, F. & CARIANI, A.  (2021) Molecular Taxonomy and Diversification of Atlantic Skates (Chondrichthyes, Rajiformes): Adding More Pieces to the Puzzle of Their Evolutionary History. Life, 11 (7): 596 https://dx.doi.org/10.3390/life11070596
DA CRUZ, V.P. & ADACHI, AMCD, RIBEIRO, G.D. & DE OLIVEIRA, P.H. & DE OLIVEIRA, C. & ORIANO, R. & DE FREITAS, R.H.A. & FORESTI, F. (2021) A shot in the dark for conservation: Evidence of illegal commerce in endemic and threatened species of elasmobranch at a public fish market in southern Brazil. Aquatic Conservation: Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/aqc.3572
DA CRUZ, V.P. & NOBILE, M.L.O. & PAIM, F.G. & ADACHI, AMCD, RIBEIRO, G.D. & FERREIRA, D.C. & PANSONATO-ALVES, J.C. & CHARVET, P. & OLIVEIRA, C. & FORESTI, F. (2021) Cytogenetic and molecular characteristics of Potamotrygon motoro and Potamotrygon sp. (Chondrichthyes, Myliobatiformes, Potamotrygonidae) from the Amazon basin: Implications for the taxonomy of the genus. Genetics and Molecular Biology, 44 (2): e20200083 https://dx.doi.org/10.1590/1678-4685-gmb-2020-0083
DE QUEIROZ, JDGR, BEZERRA, N.P.A. & MACENA, B.C.L. & HAZIN, F.H.V. (2021) Back from the dead? Not really. The tale of the Galapagos shark (Carcharhinus galapagensis) in a remote Brazilian archipelago. Biological Conservation, 256: 109097 https://dx.doi.org/10.1016/j.biocon.2021.109097
DENTI, L. & PIROLA, Y. & PREVITALI, M. & CECCATO, T. & DELLA VEDOVA, G. & RIZZI, R. & BONIZZONI, P. (2021) Shark: fishing relevant reads in an RNA-Seq sample. Bioinformatics, 37 (4): 464-472 https://dx.doi.org/10.1093/bioinformatics/btaa779
DIEZ, G. & ARREGI, L. & BASTERRETXEA, M. & CUENDE, E. & OYARZABAL, I. (2021) Preliminary observations on abundance and distribution of fish fauna in a canyon of the Bay of Biscay (ICES Division 8c). Journal of the Marine Biological Association of the United Kingdom, 101 (1): 169-178 https://dx.doi.org/10.1017/s0025315420001265
D'IGLIO, C. & SAVOCA, S. & RINELLI, P. & SPANO, N. & CAPILLO, G. (2021) Diet of the Deep-Sea Shark Galeus melastomus Rafinesque, 1810, in the Mediterranean Sea: What We Know and What We Should Know. Sustainability, 13 (7): 3962 https://dx.doi.org/10.3390/su13073962
DUREUIL, M. & AEBERHARD, W.H. & BURNETT, K.A. & HUETER, R.E. & TYMINSKI, J.P. & WORM, B. (2021) Unified natural mortality estimation for teleosts and elasmobranchs. Marine Ecology Progress Series, 667: 113–129 https://dx.doi.org/10.3354/meps13704
EBERT, D.A. & LESLIE, R.W. & WEIGMANN, S.  (2021) Etmopterus brosei sp. nov.: a new lanternshark (Squaliformes: Etmopteridae) from the southeastern Atlantic and southwestern Indian oceans, with a revised key to the Etmopterus lucifer clade. Marine Biodiversity, 51: 53 https://dx.doi.org/10.1007/s12526-021-01173-0
ELSTON, C. & COWLEY, P.D. & VON BRANDIS, R.G. & LEA, J. (2021) Residency and habitat use patterns by sympatric stingrays at a remote atoll in the Western Indian Ocean. Marine Ecology Progress Series, 662: 97-114 https://dx.doi.org/10.3354/meps13632
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FAHMI & KURNIAWAN, W. & TIBBETTS, I.R. & OKTAVIYANI, S. & DUDGEON, C.L. & BENNETT, M.B. (2021) Age and growth of the tropical oviparous shark, Chiloscyllium punctatum in Indonesian waters. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14778
FIDLER, R.Y. & ANDRADI-BROWN, D.A. & AWALUDINNOER, PADA, D. & PURWANTO, HIDAYAT, N.I. & AHMADIA, G.N. & HARBORNE, A.R. (2021) The importance of biophysical context in understanding marine protected area outcomes for coral reef fish populations. Coral Reefs, in press https://dx.doi.org/10.1007/s00338-021-02085-y
FINUCCI, B. & DUFFY, C.A.J. & BROUGH, T. & FRANCIS, M.P. & MILARDI, M. & PINKERTON, M.H. & PETERSEN, G. & STEPHENSON, F. (2021) Drivers of Spatial Distributions of Basking Shark (Cetorhinus maximus) in the Southwest Pacific. Frontiers in Marine Science, 8: 665337 https://dx.doi.org/10.3389/fmars.2021.665337
FORREST, J.A.H. & BOUCHET, P.J. & BARLEY, S.C. & MCLENNAN, A.G. & MEEUWIG, J.J. (2021) True blue: Temporal and spatial stability of pelagic wildlife at a submarine canyon. Ecosphere, 12 (3): e03423 https://dx.doi.org/10.1002/ecs2.3423
GALINDO, E. & GIRALDO, A. & NAVIA, A.F. (2021) Feeding habits and trophic interactions of four sympatric hammerhead shark species reveal trophic niche partitioning. Marine Ecology Progress Series, 665: 159-175 https://dx.doi.org/10.3354/meps13681
GALLOWAY, K.A. & PORTER, M.E. (2021) Predator-Prey Interactions Examined Using Lionfish Spine Puncture Performance.  Integrative Organismal Biology, 3 (1): obaa049 https://dx.doi.org/10.1093/iob/obaa049
GARCIA-HERAS, M. & AGUA, F. & MADIQUIDA, H. & FERNANDEZ, V.M. & DE TORRES, J. & VILLEGAS, M.A. & RUIZ-GALVEZ, M. (2021) Characterization of Glass, Shell, and Fishbone Beads on Ibo Island (Northern Mozambique) in the Context of the Indian Ocean Trade. African Archaeological Review, in press https://dx.doi.org/10.1007/s10437-021-09430-0
GERVAIS, C.R. & BROWN, C. (2021) Impact of conspecific necromones on the oxygen uptake rates of a benthic elasmobranch. Animal Behaviour, 174: 1-8 https://dx.doi.org/10.1016/j.anbehav.2021.01.009
GIOVOS, I. & SPYRIDOPOULOU, R.N.A. & DOUMPAS, N. & GLAUS, K. & KLEITOU, P. & KAZLARI, Z. & KATSADA, D. & LOUKOVITIS, D. & MANTZOUNI, I. & PAPAPETROU, M. & PAPASTAMATIOU, Y.P. & MOUTOPOULOS, D.K. (2021) Approaching the “real” state of elasmobranch fisheries and trade: A case study from the Mediterranean. Ocean & Coastal Management, 211: 105743 https://dx.doi.org/10.1016/j.ocecoaman.2021.105743
GOLANI, D. (2021) An updated Checklist of the Mediterranean fishes of Israel, with illustrations of recently recorded species and delineation of Lessepsian migrants. Zootaxa, 4956 (1): 1-108  https://dx.doi.org/10.11646/zootaxa.4956.1.1
GONZALEZ-ACOSTA, A.F. & MONSALVO-FLORES, A.E. & TOVAR-AVILA, J. & JIMENEZ-CASTANEDA, M.F. & ALEJO-PLATA, M.D. & DE LA CRUZ-AGUERO, G. (2021) Diversity and conservation of Chondrichthyes in the Gulf of California. Marine Biodiversity, 51 (3): 46  https://dx.doi.org/10.1007/s12526-021-01186-9
GRAHAM, F. (2021) Daily briefing: Glow-in-the-dark shark is biggest bioluminescent vertebrate. Nature, in press https://dx.doi.org/10.1038/d41586-021-00603-0
GRAHAM, J. & KROETZ, A.M. & POULAKIS, G.R. & SCHARER, R.M. & CARLSON, J.K. & LOWERRE-BARBIERI, S. & MORLEY, D. & REYIER, E.A. & GRUBBS, R.D. (2021) Large-scale space use of large juvenile and adult smalltooth sawfish Pristis pectinata: implications for management. Endangered Species Research, 44: 45-59 https://dx.doi.org/10.3354/esr01088
GRIFFIN, L.P. & CASSELBERRY, G.A. & HART, K.M. & JORDAAN, A. & BECKER, S.L. & NOVAK, A.J. & DEANGELIS, B.M. & POLLOCK, C.G. & LUNDGREN, I. & HILLIS-STARR, Z. & DANYLCHUK, A.J. & SKOMAL, G.B. (2021) A Novel Framework to Predict Relative Habitat Selection in Aquatic Systems: Applying Machine Learning and Resource Selection Functions to Acoustic Telemetry Data From Multiple Shark Species. Frontiers in Marine Science, 8: 631262 https://dx.doi.org/10.3389/fmars.2021.631262
GRIMWOOD, R.M. & HOLMES, E.C. & GEOGHEGAN, J.L. (2021) A Novel Rubi-Like Virus in the Pacific Electric Ray (Tetronarce californica) Reveals the Complex Evolutionary History of the Matonaviridae. Viruses-Basel, 13 (4): 585 https://dx.doi.org/10.3390/v13040585
GULAK, S.J.B. & CARLSON, J.K. (2021) Less Soak Time Saves Those upon the Line: Capture Times and Hooking Mortality of Sharks Caught on Bottom Longlines. North American Journal of Fisheries Management, in press https://dx.doi.org/10.1002/nafm.10592
HABIB, K.A. & ISLAM, M.J. (2021) Description of a new species of giant guitarfish, Glaucostegus younholeei sp. nov. (Rhinopristiformes: Glaucostegidae) from the northern Bay of Bengal, Bangladesh. Zootaxa, 4995 (1): 129–146 https://dx.doi.org/10.11646/zootaxa.4995.1.7
HAQUE, A.B. & SPAET, J.L.Y. (2021) Trade in threatened elasmobranchs in the Bay of Bengal, Bangladesh. Fisheries Research, 243: 106059 https://dx.doi.org/10.1016/j.fishres.2021.106059
HAUSER-DAVIS, R.A. & AMORIM-LOPES, C. & ARAUJO, N.L.F. & REBOUÇAS, M. & GOMES, R.A. & ROCHA, R.C.C. & DILLENBURG SAINT'PIERRE, T. & NEVES DOS SANTOS, L. (2021) On mobulid rays and metals: Metal content for the first Mobula mobular record for the state of Rio de Janeiro, Brazil and a review on metal ecotoxicology assessments for the Manta and Mobula genera. Marine Pollution Bulletin, 168: 112472 https://dx.doi.org/10.1016/j.marpolbul.2021.112472
HAYATA, M.A. & BORNATOWSKI, H. & FREITAS, R.H.A. (2021) Patterns and partitioning of food resources by elasmobranchs in southern Brazil. Environmental Biology of Fishes, in press https://dx.doi.org/10.1007/s10641-021-01082-y
HEIM, V. & DHELLEMMES, F. & SMUKALL, M.J. & GRUBER, S.H. & GUTTRIDGE, T.L. (2021) Effects of Food Provisioning on the Daily Ration and Dive Site Use of Great Hammerhead Sharks, Sphyrna mokarran. Frontiers in Marine Science, 8: 628469 https://dx.doi.org/10.3389/fmars.2021.628469
HSU, L.H. & SU, C.Y. & SUN, P.L. & CHEN, Y.L. (2021) Fusarium solani species complex infection in elasmobranchs: A case report for rough-tail stingray with valid antifungal therapy. Med Mycol Case Rep., 32: 34-38 https://dx.doi.org/10.1016/j.mmcr.2021.02.002
JAMES, K.C. & NATANSON, L.J. & FLIGHT, C. & TRIBUZIO, C. & HOEY, J. & MCCANDLESS, C. (2021) Validation of the use of vertebrae and dorsal-fin spines for age determination of spiny dogfish (Squalus acanthias) in the western North Atlantic Ocean. Fishery Bulletin, 119 (1): 41-49 https://dx.doi.org/10.7755/FB.119.1.6
JOHANSON, Z. & MANZANARES, E. & UNDERWOOD, C. & CLARK, B. & FERNANDEZ, V. & SMITH, M. (2021) Ontogenetic development of the holocephalan dentition: Morphological transitions of dentine in the absence of teeth. Journal of Anatomy, in press https://dx.doi.org/10.1111/joa.13445
JOHRI, S. & LIVINGSTON, I. & TIWARI, A. & SOLANKI, J. & BUSCH, A. & MORENO, I. & FELLOWS, S.R. & DOANE, M.P. & DINSDALE, E.A. (2021) Reducing Data Deficiencies: Preliminary Elasmobranch Fisheries Surveys in India, Identify Range Extensions and Large Proportions of Female and Juvenile Landings. Frontiers in Marine Science, 8: 619695 https://dx.doi.org/10.3389/fmars.2021.619695
JUMA, S.N. & GONG, X.X. & HU, S.J. & LV, Z.B. & SHAO, J.Z. & LIU, L.L. & CHEN, G.Q. (2021) Shark New Antigen Receptor (IgNAR): Structure, Characteristics and Potential Biomedical Applications. Cells, 10 (5): 1140 https://dx.doi.org/10.3390/cells10051140
KATSU, Y. & SHARIFUL, I.M.D. & LIN, X. & TAKAGI, W. & URUSHITANI, H. & KOHNO, S. & HYODO, S. & BAKER, M.E. (2021) N-terminal Domain Regulates Steroid Activation of Elephant Shark Glucocorticoid and Mineralocorticoid Receptors. Journal of Steroid Biochemistry and Molecular Biology, 210: 105845 https://dx.doi.org/10.1016/j.jsbmb.2021.105845
KELLER, B.A. & PUTMAN, N.F. & GRUBBS, R.D. & PORTNOY, D.S. & MURPHY, T.P. (2021) Map-like use of Earth's magnetic field in sharks. Current Biology, in press https://dx.doi.org/10.1016/j.cub.2021.03.103
KIM, S.W. & PARK, S.Y. & KWON, H. & GIRI, S.S. & KIM, S.G. & KANG, J.W. & KWON, J. & LEE, S.B. & JUNG, W.J. & LEE, J. & PARK, S.C. & KIM, J.H. (2021) Complete mitochondrial genome and phylogenetic analysis of the copper shark Carcharhinus brachyurus (Gunther, 1870). Mitochondrial DNA Part B-Resources, 6 (6): 1659-1661 https://dx.doi.org/10.1080/23802359.2021.1920863
KINA, T. & MASUDA, T. & ASAHINA, K. & SUZUKI, M. (2021) Composition of uterine milk and its changes with gestational period in red stingrays (Hemitrygon akajei). Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14716
KYNE, P.M. & OETINGER, M. & GRANT, M.I. & FEUTRY, P. (2021) Life history of the Critically Endangered largetooth sawfish: a compilation of data for population assessment and demographic modelling. Endangered Species Research, 44: 79-88 https://dx.doi.org/10.3354/esr01090
LIN, T.C. & HSIAO, W.V. & HAN, S.J. & JOUNG, S.J. & SHIAO, J.C. (2021) A direct multiplex loop-mediated isothermal amplification method to detect three CITES-listed shark species. Aquatic Conservation-Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/aqc.3592
LLOYD, C.J. & PEAKALL, J. & BURNS, A. & KEEVIL, G. & DORRELL, R. & WIGNALL, P. & FLETCHER, T. (2021) Hydrodynamic efficiency in sharks: the combined role of riblets and denticles. Bioinspiration & Biomimetics, 16: 046008 https://dx.doi.org/10.1088/1748-3190/abf3b1
LOBODA, T.S. & LASSO, C.A. & ROSA, R.S & DE CARVALHO, M.R. (2021) Two new species of freshwater stingrays of the genus Paratrygon (Chondrichthyes: Potamotrygonidae) from the Orinoco basin, with comments on the taxonomy of Paratrygon aiereba. Neotropical Ichthyology, 19 (2): e200083 https://dx.doi.org/10.1590/1982-0224-2020-0083
LOPES DA SILVA FERRETTE, B. & COELHO, R. & PEDDEMORS, V.M. & OVENDEN, J.R. & DE FRANCO, B.A. & OLIVEIRA, C. & FORESTI, F. & MENDONÇA, F.F. (2021) Global phylogeography of the smooth hammerhead shark: Glacial refugia and historical migration patterns. Aquatic Conservation: Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/aqc.3629
LUIZ MARCON, J. & DARIO MORALES-GAMBA, R. & FERNANDO MARQUES BARCELLOS, J. & LÚCIA GÓES DE ARAÚJO, M. (2021) Sex steroid hormones and the associated morphological changes in the reproductive tract of free-living males of the cururu stingray Potamotrygon wallacei. General and Comparative Endocrinology, 309: 113786 https://dx.doi.org/10.1016/j.ygcen.2021.113786
LYONS, K. & ADAMS, D.H. & BIZZARRO, J.J. (2021) Evaluation of muscle tissue as a non-lethal proxy for liver and brain organic contaminant loads in an elasmobranch, the Bonnethead Shark (Sphyrna tiburo). Mar Pollut Bull., 167: 112327 https://dx.doi.org/10.1016/j.marpolbul.2021.112327
MATICH, P. & SHIPLEY, O.N. & WEIDELI, O.C. (2021) Quantifying spatial variation in isotopic baselines reveals size-based feeding in a model estuarine predator: implications for trophic studies in dynamic ecotones. Marine Biology, 168: 108 https://dx.doi.org/10.1007/s00227-021-03920-0
MAURICE, L. & LE CROIZIER, G. & MORALES, G. & CARPINTERO, N. & GUAYASAMIN, J.M. & SONKE, J. & PAEZ-ROSAS, D. & POINT, D. & BUSTOS, W. & OCHOA-HERRERA, V. (2021) Concentrations and stable isotopes of mercury in sharks of the Galapagos Marine Reserve: Human health concerns and feeding patterns. Ecotoxicology and Environmental Safety, 215: 112122 https://dx.doi.org/10.1016/j.ecoenv.2021.112122
MILANKOVIC, H.R. & RAY, N.D. & GENTLE, L.K. & KRUGER, C. & JACOBS, E. & FERREIRA, C.J. (2021) Seasonal occurrence and sexual segregation of great white sharks Carcharodon carcharias in Mossel Bay, South Africa. Environmental Biology of Fishes, 104 (5): 555-568 https://dx.doi.org/10.1007/s10641-021-01094-8
MOLINA-SALGADO, P. & ALFARO-SHIGUETO, J. & GONZÁLEZ-PESTANA, A. (2021) Diet of the rasptail skate, Rostroraja velezi (Rajiformes: Rajidae), off Piura, Peru. Ciencias Marinas, 47 (2): 127–138 https://dx.doi.org/10.7773/cm.v47i2.3132
MORGAN, D.L. & LEAR, K.O. & DOBINSON, E. & GLEISS, A.C. & FAZELDEAN, T. & PILLANS, R.D. & BEATTY, S.J. & WHITTY, J.M. (2021) Seasonal use of a macrotidal estuary by the endangered dwarf sawfish, Pristis clavata. Aquatic Conservation-Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/aqc.3578
MOURIER, J. & PLANES, S. (2021) Kinship does not predict the structure of a shark social network. Behavioral Ecology, 32 (2): 211-222 https://dx.doi.org/10.1093/beheco/araa119
OLIVEIRA, C.D.L. & OLIVEIRA, C.Y.B. & CAMILO, J.P.G. & BATISTA, V.D. (2021) Demographic analysis reveals a population decline of the Longnose stingray Hypanus guttatus in Northeastern Brazil. Regional Studies in Marine Science, 41: 101554 https://dx.doi.org/10.1016/j.rsma.2020.101554
ORTEGA, R.P. (2021) Ethical controversy swirls around shark fossil from Mexico. Science, 372 (6540): 332-333 https://dx.doi.org/10.1126/science.372.6540.332
PADILHA, F.L.A. & COLBACHINI, H. & RAMOS, S.D. & REISFELD, L.C. & HENRIQUE, P.C. & LEITE, R.F. & LOSANO, J.D.D. & NICHI, M. & SOUSA, R.G.B. & GUTIERREZ, R.C. & JORGE-NETO, P.N. & PIZZUTTO, C.S. (2021) Validation of manual semen collection methods and sperm evaluation in living freshwater stingrays (Potamotrygon falkneri) kept in ex situ conditions. Environmental Biology of Fishes, in press https://dx.doi.org/10.1007/s10641-021-01086-8
PAEZ-ROSAS, D. & SUAREZ-MONCADA, J. & ELORRIAGA-VERPLANCKEN, F.R. & PROANO, A. & ARNES-URGELLES, C. & SALINAS-DE-LEON, P. & GALVAN-MAGANA, F. (2021) Trophic variation during the early stages of blacktip sharks (Carcharhinus limbatus) within coastal nurseries of the Galapagos Marine Reserve. Journal of Sea Research, 170: 102023 https://dx.doi.org/10.1016/j.seares.2021.102023
PANCALDI, F. & PAEZ-OSUNA, F. & MARMOLEJO-RODRIGUEZ, A.J. & WHITEHEAD, D.A. & GONZALEZ-ARMAS, R. & SOTO-JIMENEZ, M.F. & O'HARA, T. & VAZQUEZ-HAIKIN, A. & GALVAN-MAGANA, F. (2021) Variation of essential and non-essential trace elements in whale shark epidermis associated to two different feeding areas of the Gulf of California. Environmental Science and Pollution Research, in press https://dx.doi.org/10.1007/s11356-021-13364-0
PANCALDI, F. & PAEZ-OSUNA, F. & SOTO-JIMENEZ, M.F. & WHITEHEAD, D.A. & GONZALEZ-ARMAS, R. & VAZQUEZ-HAIKIN, A. & BECERRIL-GARCIA, E.E. & GALVAN-MAGANA, F. (2021) Concentrations of Silver, Chrome, Manganese and Nickel in Two Stranded Whale Sharks (Rhincodon typus) from the Gulf of California. Bulletin of Environmental Contamination and Toxicology, in press https://dx.doi.org/10.1007/s00128-021-03244-1
PEREZ-ROJAS, J.G. & MEJIA-FALLA, P.A. & NAVIA, A.F. & TARAZONA, A.M. & PARDO-CARRASCO, S.C. (2021) Hematology and blood biochemistry profile of the freshwater stingray Potamotrygon magdalenae as a tool for population assessment in artificial environments. Brazilian Journal of Biology, 82: e233780 https://dx.doi.org/10.1590/1519-6984.233780
PIMIENTO, C. & PYENSON, N.D. (2021) When sharks nearly disappeared. Science, 372 (6546): 1036-1037 https://dx.doi.org/10.1126/science.abj2088
PINI-FITZSIMMONS, J. & KNOTT, N.A. & BROWN, C. (2021) Heterarchy Reveals Social Organization of a Smooth Stingray (Bathytoshia brevicaudata) Population in a Provisioned Food Context. Frontiers in Marine Science, 8: 641761 https://dx.doi.org/10.3389/fmars.2021.641761
REZAIE-ATAGHOLIPOUR, M. & JABADO, R.W. & OWFI, F. & HESNI, M.A. & EBERT, D.A. (2021) Lost and found: rediscovery of the extinct tentacled butterfly ray Gymnura tentaculata in Iranian waters. Oryx, 55 (4): 489-490 https://dx.doi.org/10.1017/S0030605321000521
ROUSSOS, I. & MEGALOFONOU, P. (2021) Ontogenetic shifts and sexual dimorphism in the brain organization of the small-spotted catshark Scyliorhinus canicula. Journal of the Marine Biological Association of the United Kingdom, 101 (1): 189-198 https://dx.doi.org/10.1017/s0025315420001368
SANTOS DOS SANTOS, P.R. (2021) Recreational fishing as a source for the monitoring of a critically endangered shark in southern Brazil. Fisheries Research, 241: 106006 https://dx.doi.org/10.1016/j.fishres.2021.106006
SCHLUESSEL, V. & RICK, I.P. & SEIFERT, F.D. & BAUMANN, C. & DAVIES, W.I.L. (2021) Not just shades of grey: life is full of colour for the ocellate freshwater river stingray (Potamotrygon motoro). Journal of Experimental Biology, 224 (9): jeb226142 https://dx.doi.org/10.1242/jeb.226142
SCHOEN, A.N. & TREBERG, J.R. & WHEATON, C.J. & MYLNICZENKO, N. & GARY, ANDERSON, W. (2021) Energy and corticosteroid mobilization following an induced stress response in an elasmobranch fish, the North Pacific spiny dogfish (Squalus acanthias suckleyi). General and Comparative Endocrinology,  310: 113799 https://dx.doi.org/10.1016/j.ygcen.2021.113799
SEGURA-COBENA, E. & ALFARO-SHIGUETO, J. & MANGEL, J. & URZUA, A. & GORSKI, K. (2021) Stable isotope and fatty acid analyses reveal significant differences in trophic niches of smooth hammerhead Sphyrna zygaena (Carcharhiniformes) among three nursery areas in northern Humboldt Current System. Peerj, 9: e11283 https://dx.doi.org/10.7717/peerj.11283
SHELLEM, C.T. & ELLIS, J.I. & COKER, D.J. & BERUMEN, M.L. (2021) Red Sea fish market assessments indicate high species diversity and potential overexploitation. Fisheries Research, 239 https://dx.doi.org/10.1016/j.fishres.2021.105922
SOMSAP, N. & SRAKAEW, N. & CHATCHAVALVANICH, K. (2021) Microanatomy of the female reproductive system of the viviparous freshwater whipray Fluvitrygon signifer (Chondrichthyes: Myliobatiformes: Dasyatidae). II. The genital duct. BMC Zoology, 6 (1): 11 https://dx.doi.org/10.1186/s40850-021-00070-0
SOYKAN, O. & KINACIGIL, H.T. (2021) Length-Weight Relationship of some Discarded Fish Species with Emphasis on Length at Maturity from the Central Aegean Sea, Turkey. Thalassas, in press https://dx.doi.org/10.1007/s41208-021-00291-7
STRAUBE, N. & LYRA, M.L. & PAIJMANS, J.L.A. & PREICK, M. & BASLER, N. & PENNER, J. & RÖDEL, M.-O. & WESTBURY, M.V. & HADDAD, C.F.B. & BARLOW, A. & HOFREITER, M. (2021) Successful application of ancient DNA extraction and library construction protocols to museum wet collection specimens. Molecular Ecology Resources, in press https://dx.doi.org/10.1111/1755-0998.13433
SUMIKAWA, H. & MIYOSHI, T. (2021) The pressure drag reduction effect of tandem swimming by Caranx sexfasciatus and Rhincodon typus. Ichthyological Research, in press https://dx.doi.org/10.1007/s10228-021-00822-5
TANNA, A. & FERNANDO, D. & GOBIRAJ, R. & PATHIRANA, B.M. & THILAKARATNA, S. & JABADO, R.W. (2021) Where have all the sawfishes gone? Perspectives on declines of these Critically Endangered species in Sri Lanka. Aquatic Conservation: Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/aqc.3617
TERRAPON, M. & KISZKA, J.J. & WAGNER, J. (2021) Observations of Killer Whale (Orcinus orca) Feeding Behavior in the Tropical Waters of the Northern Mozambique Channel Island of Mayotte, Southwest Indian Ocean. Aquatic Mammals, 47 (2): 196-205 https://dx.doi.org/10.1578/am.47.2.2021.196
TORRES-HUERTA, A.M. & CRUZ-MARTINEZ, A. & RAMIREZ-CHAVEZ, E.J. (2021) Spatial pattern of distribution and reproductive strategy of vermiculate electric-rays Narcine vermiculatus. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14735
VERKAMP, H.J. & SKOMAL, G. & WINTON, M. & SULIKOWSKI, J.A. (2021) Using reproductive hormone concentrations from the muscle of white sharks Carcharodon carcharias to evaluate reproductive status in the Northwest Atlantic Ocean. Endangered Species Research, 44: 231-236 https://dx.doi.org/10.3354/esr01109
VIJAYAN, D.K. & RAMAN, S.P. & DARA, P.K. & JACOB, R.M. & MATHEW, S. & RANGASAMY, A. & NAGARAJARAO, R.C. (2021) In vivo anti-lipidemic and antioxidant potential of collagen peptides obtained from great hammerhead shark skin waste. Journal of Food Science and Technology-Mysore, in press https://dx.doi.org/10.1007/s13197-021-05118-0
WALKER, T.I. & DAY, R.W. & AWRUCH, C.A. & BELL, J.D. & BRACCINI, J.M. & DAPP, D.R. & FINOTTO, L. & FRICK, L.H. & GARCES-GARCIA, K.C. & GUIDA, L. & HUVENEERS, C. & MARTINS, C.L. & ROCHOWSKI, B.E.A. & TOVAR-AVILA, J. & TRINNIE, F.I. & REINA, R.D. (2021) Ecological vulnerability of the chondrichthyan fauna of southern Australia to the stressors of climate change, fishing and other anthropogenic hazards. Fish and Fisheries, in press https://dx.doi.org/10.1111/faf.12571
WALLACE, S.S. (2021) Sharks, Skates, Rays and Chimeras of British Columbia. Fisheries, in press https://dx.doi.org/10.1002/fsh.10611
WANG, J.Q. & GAO, C.X. & WU, F. & GAO, X.D. & CHEN, J. & DAI, X.J. & TIAN, S.Q. & CHEN, Y. (2021) The discards and bycatch of Chinese tuna longline fleets in the Pacific Ocean from 2010 to 2018. Biological Conservation, 255: 109011 https://dx.doi.org/10.1016/j.biocon.2021.109011
WAXMAN, L. & MYLNICZENKO, N.D. & FUSTUKJIAN, A. & STACY, N.I. (2021) What is your diagnosis? Reproductive tract fluid from a radiated tortoise (Astrochelys radiata) and southern stingray (Hypanus americanus). Veterinary Clinical Pathology, in press https://dx.doi.org/10.1111/vcp.12937
WEIGMANN, S. & EBERT, D.A. & SÉRET, B.  (2021) Resolution of the Acroteriobatus leucospilus species complex, with a redescription of A. leucospilus (Norman, 1926) and descriptions of two new western Indian Ocean species of Acroteriobatus (Rhinopristiformes, Rhinobatidae). Marine Biodiversity, 51: 58 https://dx.doi.org/10.1007/s12526-021-01208-6
WEINRAUCH, A.M. & HOOGENBOOM, J.L. & ANDERSON, W.G. (2021) A review of reductionist methods in fish gastrointestinal tract physiology. Comparative Biochemistry and Physiology B-Biochemistry & Molecular Biology, 254: 110571 https://dx.doi.org/10.1016/j.cbpb.2021.110571
WEND, C.M. & GOOLSBY, C. & SCHULER, K. & FISCHER, ST. & LEVY, M.J. (2021) Tourniquet Use in Animal Attacks: An Analysis of News Media Reports. Cureus, 13 (3): e13926 https://dx.doi.org/10.7759/cureus.13926
WHITE, W.T. & FRICKE, R. (2021) Raja mauritaniensis: a replacement name for Raja africana Capapé, 1977 (Rajiformes: Rajidae), a junior homonym of Raja africana Bloch & Schneider, 1801 (Myliobatiformes: Dasyatidae). Zootaxa, 4970 (2): 399–400 https://dx.doi.org/10.11646/zootaxa.4970.2.12
WILMERS, J. & WALDRON, M. & BARGMANN, S. (2021) Hierarchical Microstructure of Tooth Enameloid in Two Lamniform Shark Species, Carcharias taurus and Isurus oxyrinchus. Nanomaterials, 11 (4): 969 https://dx.doi.org/10.3390/nano11040969
WYFFELS, J.T. & ADAMS, L.M. & BULMAN, F. & FUSTUKJIAN, A. & HYATT, M.W. & FELDHEIM, K.A. & PENFOLD, L.M. (2021) Artificial insemination and parthenogenesis in the whitespotted bamboo shark Chiloscyllium plagiosum. Scientific Reports, 11 (1): 9966 https://dx.doi.org/10.1038/s41598-021-88568-y
ZEA-DE LA CRUZ, H. & TOVAR-AVILA, J. & MEINERS-MANDUJANO, C. & JIMENEZ-BADILLO, L. & OVIEDO-PEREZ, J.L. (2021) Determining potential management strategies for the elasmobranchs bycatch of the Mexican shrimp trawl fishery of the Gulf of Mexico through a vulnerability analysis. Regional Studies in Marine Science, 42: 101626 https://dx.doi.org/10.1016/j.rsma.2021.101626

Extinct Chondrichthyes:

ADNET, S. & FEICHTINGER, I. & HARZHAUSER, M. & POLLERSPÖCK, J. (2021) A mesopelagic selachian fauna from the middle Eocene of St. Pankraz (Austria) reveals homogeneity in deep-marine environments during the warm period in Europe. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, 301 (1): 25-63  https://dx.doi.org/10.1127/njgpa/2021/0996
AGNOLIN, F.L. & BOGAN, S. & CASADIO, S. (2021) First marine ichthyofauna from the late Eocene of Santa Cruz province, patagonia, Argentina. Journal of South American Earth Sciences, 107: 103064 https://dx.doi.org/10.1016/j.jsames.2020.103064
BENNETT, C.E. & KEARSEY, T.I. & DAVIES, S.J. & LENG, M.J. & MILLWARD, D. & SMITHSON, T.R. & BRAND, P.J. & BROWNE, M.A.E. & CARPENTER, D.K. & MARSHALL, J.E.A. & DULSON, H. & CURRY, L. (2021) Palaeoecology and palaeoenvironment of Mississippian coastal lakes and marshes during the early terrestrialisation of tetrapods. Palaeogeography Palaeoclimatology Palaeoecology, 564: 110194 https://dx.doi.org/10.1016/j.palaeo.2020.110194
CEBALLOS-IZQUIERDO, Y. & VINOLA-LOPEZ, L.W. & BORGES-SELLEN, C.R. & ARANO-RUIZ, A.F. (2021) Late Cretaceous sharks from Cuba, first record of Serratolamna serrata (Agassiz) (Lamniformes, Serratolamnidae). Geobios, 65: 1-6 https://dx.doi.org/10.1016/j.geobios.2021.01.002    
COLLARETA, A. & LANDINI, W. & BIANUCCI, G. & DI CELMA, C. (2021) Until Panama do us part: new finds from the Pliocene of Ecuador provide insights into the origin and palaeobiogeographic history of the extant requiem sharks Carcharhinus acronotus and Nasolamia velox. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 300 (1): 103-115  https://dx.doi.org/10.1127/njgpa/2021/0981    
COLLARETA, A. & MARX, F.G. & CASATI, S. & DI CENCIO, A. & MERELLA, M. & BIANUCCI, G. (2021) A cetotheriid whale from the upper Miocene of the Mediterranean. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, 301 (1):  9-16  https://dx.doi.org/10.1127/njgpa/2021/0994    
COLLARETA, A. & MERELLA, M. & CASATI, S. & DI CENCIO, A. (2021) First fossils of the extant blacktip shark Carcharhinus limbatus from Europe and the Mediterranean Basin. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, 301 (1):  109-118  https://dx.doi.org/10.1127/njgpa/2021/1002    
DANKINA, D. & SPIRIDONOV, A. & RACZYŃSKI, P. & RADZEVIČIUS, S. (2021) Late Permian ichthyofauna from the North-Sudetic Basin, SW Poland. Acta Palaeontologica Polonica, 66: in press  https://dx.doi.org/10.4202/app.00839.2020    
DILLON, E.M. & MCCAULEY,  D.J. & MORALES-SALDAÑA, J.M. & LEONARD,  N.D. & ZHAO, J.-X. & O’DEA, A. (2021) Fossil dermal denticles reveal the preexploitation baseline of a Caribbean coral reef shark community. Proceedings of the National Academy of Sciences of the United States of America, 118 (29): e2017735118 https://dx.doi.org/10.1073/pnas.2017735118
FEICHTINGER, I. & IVANOV, A.O. & WINKLER, V. & DOJEN, C. & KINDLIMANN, R. & KRIWET, J. & PFAFF, C. & SCHRAUT, G. & STUMPF, S. (2021) Scarce ctenacanthiform sharks from the Mississippian of Austria with an analysis of Carboniferous elasmobranch diversity in response to climatic and environmental changes. Journal of Vertebrate Paleontology, in press https://dx.doi.org/10.1080/02724634.2021.1925902
FILEK, T. & HOFMAYER, F. & FEICHTINGER, I. & BERNING, B. & POLLERSPÖCK, J. & ZWICKER, J. & SMRZKA, D. & PECKMANN, J. & KRANNER, M. & MANDIC, O. & REICHENBACHER, B. & KROH, A. & UCHMAN, A. & ROETZEL, R. & HARZHAUSER, M. (2021) Environmental conditions during the late Oligocene transgression in the North Alpine Foreland Basin (Eferding Formation, Egerian) – a multidisciplinary approach. Palaeogeography, Palaeoclimatology, Palaeoecology, in press https://dx.doi.org/10.1016/j.palaeo.2021.110527    
KLUG, C. & SCHWEIGERT, G. & HOFFMANN, R. & WEIS, R. & DE BAETS, K. (2021) Fossilized leftover falls as sources of palaeoecological data: a ‘pabulite’ comprising a crustacean, a belemnite and a vertebrate from the Early Jurassic Posidonia Shale. Swiss Journal of Palaeontology, 140: 10 https://dx.doi.org/10.1186/s13358-021-00225-z    
KOVALCHUK, O. & BARKASZI, Z. (2021) Oligocene basking sharks (Lamniformes, Cetorhinidae) of the Carpathian Basin with a reconsideration of the role of gill rakers in species diagnostics. Journal of Vertebrate Paleontology, in press https://dx.doi.org/10.1080/02724634.2021.1929269    
LUCCISANO, V. & PRADEL, A. & AMIOT, R. & GAND, G. & STEYER, J.-S. & CUNY, G. (2021) A new Triodus shark species (Xenacanthidae, Xenacanthiformes) from the lowermost Permian of France and its paleobiogeographic implications. Journal of Vertebrate Paleontology, in press https://dx.doi.org/10.1080/02724634.2021.1926470    
RANA, R.S. & PATEL, R. & CICIMURRI, D.J. & EBERSOLE, J.A. (2021) Additions to the elasmobranch assemblage from the Bandah Formation (middle Eocene, Bartonian), Jaisalmer District, Rajasthan, India, and the palaeobiogeographic implications of the fauna. Palaeovertebrata 44 (2): e1 https://dx.doi.org/10.18563/pv.44.2.e1    
SCHLESIGER, E. & KOGAN, I. & FISCHER, J. (2021) Isolated Elasmobranch Vertebrae from the Cretaceous of Saxony: Taxonomic and Palaeobiological Evaluation. Abstract. In: XVIII Conference of the European Association of Vertebrate Palaeontologists Online, 5 9 July 2021: 147      
SIBERT, E.C. & RUBIN, L.D. (2021) An early Miocene extinction in pelagic sharks Science, 372 (6546): 1105-1107 https://dx.doi.org/10.1126/science.aaz3549    
STUMPF, S. & ETCHES, S. & UNDERWOOD, C.J. & KRIWET, J. (2021) Durnonovariaodus maiseyi gen. et sp. nov., a new hybodontiform shark-like chondrichthyan from the Upper Jurassic Kimmeridge Clay Formation of England. Peerj, 9: e11362 https://dx.doi.org/10.7717/peerj.11362    
TAKAKUWA, Y. & KIMURA, T. & HASEGAWA, Y. (2021) Fossil sharks from the lower Pleistocene Komiya Formation, Kazusa Group, Tokyo, Japan: The teeth associated with the skeleton of Eschrichtius akishimaensis. Bulletin of Gunma Museum of Natural History, 25: 49-58
YUN, C.-G, (2021) First Deep-Sea Shark Fossil Teeth from the Miocene of South Korea. Zoodiversity, 55 (3): 225–232  https://dx.doi.org/10.15407/zoo2021.03.225    

BEVERIDGE, I. & KOEHLER, A. & APPY, R.G. (2021) Eutetrarhynchus pacificus n. sp. (Cestoda: Trypanorhyncha) from Raja inornata Jordan & Gilbert (Batoidea: Rajiformes) off the coast of California with comments on congeners. Systematic Parasitology, in press https://dx.doi.org/10.1007/s11230-021-09978-0
HERMANS, R. & VANHOVE, M.P.M. & DITRICH, O. & TYML, T. & GELNAR, M. & ARTOIS, T. & KMENTOVA, N. (2021) Parasitic flatworms infecting thorny skate, Amblyraja radiata: Infection by the monogeneans Acanthocotyle verrilli and Rajonchocotyle emarginata in Svalbard. Parasitology International, 81  https://dx.doi.org/10.1016/j.parint.2020.102261



Unexpected presence of great white sharks in Gulf of California

Date: April 21, 2021
Source: University of Delaware
Summary: A new study suggests the white shark population for the eastern north Pacific, especially those listed in the Gulf of California, might be underestimated. Researchers found that the mortality rates for these white sharks might be underestimated as well, as an illicit fishery for the species was uncovered in the Gulf of California, suggesting that fishers were killing many more white sharks than has been previously understood.

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Largest-ever study of artificial insemination in sharks -- and the occasional 'virgin birth'

Date: May 13, 2021
Source: Field Museum
Summary: Scientists help protect sharks by developing aquarium breeding programs that pair up individuals in ways that increase genetic diversity. In a new study, scientists undertook the largest-ever effort to artificially inseminate sharks. Their work resulted in 97 new baby sharks, including ones whose parents live on opposite sides of the country and a few that don't have fathers at all.

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White shark population is small but healthy off the coast of Central California

Date: May 19, 2021
Source: Oregon State University
Summary: The population of white sharks that call the Central California coast their primary home is holding steady at about 300 animals and shows some signs of growth, a new long-term study of the species has shown.

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3,000-year-old shark attack victim

Date: June 23, 2021
Source: University of Oxford
Summary: Researchers reveal their discovery of a 3,000-year-old victim - attacked by a shark in the Seto Inland Sea of the Japanese archipelago.

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Fossil shark scales provide a glimpse of reef predator populations before human impact

Date: July 5, 2021
Source: University of California - Santa Barbara
Summary: Scientists recently made news by using fossil shark scales to reconstruct shark communities from millions of years ago.
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Why are some fish warm-blooded? Predatory sharks gain speed advantage, study finds

Date: July 1, 2021
Source: Trinity College Dublin
Summary: New research from marine biologists offers answers to a fundamental puzzle that had until now remained unsolved: why are some fish warm-blooded when most are not? It turns out that while (warm-blooded) fish able to regulate their own body temperatures can swim faster, they do not live in waters spanning a broader range of temperatures.

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