NEWSLETTER 07/2020 06.07.2020

Please acknowledge use of the database www.shark-references.com in your publications, and cite: 

Pollerspöck, J. & Straube, N. 2020, Bibliography database of living/fossil sharks, rays and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali), www.shark-references.com, World Wide Web electronic publication, Version 2020

New project at shark-references!

for more information check the pdf or video

new images of Oxynotus bruniensis (OGILBY, 1893) for our project Toothmorphology!

and a new paper (co-authored by Jürgen team shark-references) is just out

free download via 

New book/free download from our partner Hakan Kabasakal

A Field Guide to the Sharks of Turkish Waters.
Turkish Marine Research Foundation (TUDAV) Publication No: 55, Istanbul, Turkey. 133 p.


Would you like to become a shark-reference partner? Please contanct us per E-mail!

Partner in Google-Maps




New Images

Many thanks to the following people for providing images:

Frederik Mollen (Elasmobranch Research Belgium) for the images of Neoraja caerulea (STEHMANN, 1976)

L. Ignacio Contreras, Laboratorio de Zoología de Vertebrados, Facultad de Ciencias, U. de Chile, for images of

Roxanie Naasan Agah, Nikos Doumpas and Ioannis Giovos, www.isea.com.gr for a image of Oxynotus centrina (LINNAEUS, 1758) and Dipturus oxyrinchus (LINNAEUS, 1758)

Gerrica Hightower, Australia for a image of Trygonoptera testacea MÜLLER & HENLE, 1841

Charlie Underwood, Natural History Museum, London for images of Planohybodus peterboroughensis REES & UNDERWOOD, 2008

Thomas Reinecke, Bochum for image of 
Raja thiedei REINECKE, 2015
Oligoraja pristina REINECKE, 2015
Torpedo chattica REINECKE, 2015
Rhynchobatus vincenti JAEKEL, 1894
Rhinobatos bruxelliensis (JAEKEL, 1894)
Carcharoides catticus (PHILIPPI, 1846)
Carcharoides totuserratus AMEGHINO, 1901
Carcharoides lipsiensis REINECKE, VON DER HOCHT, GILLE & KINDLIMANN, 2018
Pachyscyllium braaschi REINECKE, MOTHS, GRANT & BREITKREUZ, 2005
Pachyscyllium albigensis REINECKE, MOTHS, GRANT & BREITKREUZ, 2005
Scyliorhinus venloensis WEILER, 1943

Jean-Francois LHOMME (www.vertebres-fossiles.com) for images of 
Jaekelotodus robustus (LERICHE, 1922)
Onchopristis numidus (HAUG, 1905)
Sylvestrilamia teretidens (WHITE, 1931)
Squatina prima (WINKLER, 1874)

Giuseppe Marramà, Università degli Studi di Torino, Italy for images of 
Eogaleus bolcensis Cappetta, 1975
Titanonarke molini (JAEKEL, 1894)
Narcine molini JAEKEL, 1894 valid as Titanonarke molini (JAEKEL, 1894)
Lessiniabatis aenigmatica MARRAMÀ, CARNEVALE, GIUSBERTI, NAYLOR & KRIWET, 2019

Christopher Duffin for a image of Recurvacanthus uniserialis DUFFIN, 1981

John Maisey for a image of Cretasquatina americana MAISEY, EHRET & DENTON, 2020

Missing papers:

Many thanks to all friends of shark-references, who sent us some missing papers last month!

Shark-References would kindly like to ask you for your contribution to this project.

At the moment we are looking for some of the following papers:

Extinct Chondrichthyes:
NEWTON, E.T.  (1878) The chimeroid fishes of the British Cretaceous rocks. Memoirs of the Geological Survey of the United Kingdom, Monograph, 4: 1–64

CHAPMAN, F.  (1918) Descriptions and revisions of the Cretaceous and Tertiary fish-remains of New-Zealand. New Zealand Department of mines, Geological Survey Branch, Palaeontological Bulletin, 7: 1–45, pl. 1–9

MATSUMOTO, H. (1936) Upper Miocene vertebrates from Kumanodô, Natori district, province of Rikuzen. Dobutsugaku Zasshi, 48: 475–480, 5 fig.

ALVINERIE, J. & ANDREIEFF, P. & ANGLADA, R. & AUBERT, J. & CAPPETTA, H. & CARALP, M. & CARATINI, C. & CARBONNEL, G. & CATZIGRAS, F. & COURME-RAULT, M.-D. & CHATEAUNEUF, J.-J. & DEMARCQ, G. & DUCASSE, O. & FATTON, E. & GLAÇON, G. & LABRACHERIE, M. & LAURIAT, A. & LE CALVEZ, Y. & LORENZ, C. & MAGNE, J. & MARGEREL, J.-P. & POIGNANT, A. & PUJOL, C. & ROGER, J. & ROMAN, J. & BLONDEAU, A. & MULLER, C. (1973) A propos de la limite oligo-miocène: résultats préliminaires d'une recherche collective sur les gisements d'Escornébéou (Saint-Géours-de-Maremne, Landes, Aquitaine méridionale). Présence de Globigerinoides dans les faunes de l'Oligocène supérieur. Comptes rendus sommaires des séances de la Société géologique de France: 75–76

BOYD, B.M. (2016) Fossil sharks and rays of Gainesville creeks; Alachua County, Florida: Hogtown group; (middle Miocene to lower Pliocene). Florida Paleontological Society, Special Publication

Extant Chondrichthyes:

KAMOHARA, T. (1943) Some unrecorded and two new fishes from Prov. Tosa, Japan. Bulletin of the Biogeographical Society of Japan, 13 (17): 125–137

DE BUEN, F. (1950) Contribuciones a la Ictiología. II. El tiburón vitamínico de la costa uruguaya Galeorhinus vitaminicus nov. sp., y algunas consideraciones generales sobre su biología. Publicaciones Cientificas, Servicio Oceanografico y de Pesca, Ministerio de Industrias y Trabajo, Montevideo No. 4: 153–162.

CADENAT, J. (1951) Initiations Africaines. III. Poissons de Mer du Sénégal. Institute Francais d'Afrique Noire. Initiations Africaines. III. Poissons de Mer du Sénégal.: 1–345

WEIBEZAHN, F.H. (1953) Una nueva especie de Scyliorhinus de Venezuela (Chondrichthyes - Elasmobranchii). Novedades cientificas. Serie zoológica. Museo de Historia Natural La Salle, 9: 1–7.

SMITH, J.L.B. (1958) The mystery killer, the new shark Carcharhinus vanrooyeni. Veld & Vlei, 3 (9): 12–14, 28.

SICCARDI, E. (1961) Cetorhinus en el Atlantico sur (Elasmobranchii: Cetorhinidae). Actas y trabajos del Primer Congreso Sudamericano de Zoologia, 4 (5): 251–263

GUBANOV, E.P. & SCHLEIB, N.A. (1980) Sharks of the Arabian Gulf. Kuwait Ministry of Public Works, Agracultural Department, Fisheries Division. Sharks of the Arabian Gulf.: 1–69

DOLGANOV, V.N. (1983) Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod. [Manual for identification of cartilaginous fishes of Far East seas of USSR and adjacent waters.] TINRO, Vladivostok. Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod.: 92 pp.

Please support www.shark-references.com and send missing papers (not listed papers or papers without the info-symbol) to juergen.pollerspoeck@shark-references.com or nicolas.straube@shark-references.com



Upcoming Meetings:



IMCC6 to be held online

In light of the challenges the global community faces from the covid-19 pandemic, we have decided change from an in-person meeting to an online IMCC6.


The 6th International Marine Conservation Congress

To conserve the world’s oceans we must go beyond science and use it to inform policy and management to catalyze change. The International Marine Conservation Congress (IMCC) brings together conservation professionals and students to develop new and powerful tools to further marine conservation science and policy.

With over 700 marine conservation professionals and students in attendance, IMCC is the most important international event for anyone involved or interested in marine conservation. The meeting brings together marine conservationists from many walks of life including but not limited to scientists, practitioners, educators, policy-makers, artists and journalists.

IMCC6 takes place in the sailing city of Kiel, Germany. Join us in Kiel as we come together to help Make Marine Science Matter!

for more information please visit https://conbio.org/mini-sites/imcc6

Puebla, México, 19 - 23/04/2021

The Sociedad Mexicana de Peces Cartilaginosos, A. C. (SOMEPEC), is a non-profit organization that organizes the IX National Symposium of Sharks and Rays, and II Latin American Congress Sharks, Rays and Chimeras.

La Sociedad Mexicana de Peces Cartilaginosos, A. C. (SOMEPEC), es una organización sin fines de lucro, que busca promover el estudio científico de los tiburones y rayas, así como uso racional. Fiel a su objetivo de crear espacios para el intercambio de experiencias y avances de las diferentes líneas de investigación sobre tiburones y rayas, que se desarrollan en México y el resto del mundo, organiza el IX Simposium Nacional de Tiburones y Rayas, y II Congreso Latinoamericano de Tiburones, Rayas y Quimeras.




Eighth International Meeting on the enhancement and preservation of paleontological heritage, at the science faculty of El Jadida, Morocco from November 25 to 27, 2020 (visit the web page : http://www.fs.ucd.ac.ma/fs/riv3p8).



WCMB 2020

We are excited to announce that the WCMB will be going ahead as planned in Auckland, but we will also be offering an online version of the conference for those that are not able to attend in person. Travel restrictions within New Zealand have been lifted so we will be partnering with the New Zealand Marine Sciences Society (who had to cancel their 2020 annual conference) as a means of increasing local participation in the conference. To accommodate our new partners the abstract submission deadline has been extended until the 14th June 2020.

Key Dates

• Early-bird Registration closes: 1 September 2020
• Late Registration starts: 1 December 2020
• Conference commences: 14 December 2020


Extant Chondrichthyes:

 (2020): Squalus shiraii sp. nov. (Squaliformes, Squalidae), a new species of dogfish shark from Japan with regional nominal species revisited. Zoosystematics and Evolution, 96 (2): 275-311
New species: Squalus shiraii
Abstract: A new species of deep-water dogfish shark, Squalus shiraii sp. nov., is described herein as endemic to the tropical waters off Southern Japan. This species has been largely misidentified with S. mitsukurii. However, morphological, meristic and morphometric evidence support it to be a separate and undescribed species. Squalus shiraii sp. nov. differs from this species by having body brown in colour dorsally, caudal fin with ventral and dorsal tips markedly tapered and broadly white, dermal denticles uniscuspidate and lanceolate and larger number of precaudal (91–94) and total vertebrae (120–123) (vs. body dark grey to black; caudal fin with ventral and dorsal tips rounded and not white in colour; denticles tricuspidate and rhomboid; 86–90 precaudal and 116–117 total vertebrae). Squalus shiraii sp. nov. is also clearly separated from other Japanese congeners which are herein revisited to include six species, based on the examination of over 150 specimens caught from Japanese waters that were available in ichthyological collections: S. mitsukuriiS. japonicusS. acutirostrisS. brevirostris and S. suckleyiSqualus mitsukuriiS. japonicus and S. brevirostris are re-described in detail and the neotype of S. japonicus is herein designated. Squalus acutirostris is treated as a valid species with occurrences in Japan, China and Taiwan and, thus, a provisional diagnosis is given, as well as an updated diagnosis of S. suckleyi. A key to Squalus species from the North-western Pacific Ocean is given and main morphological differences between S. shiraii sp. nov. and the closest related species are discussed.

Extinct Chondrichthyes:

 (2020): A New Genus of Late Cretaceous Angel Shark (Elasmobranchii; Squatinidae), with Comments on Squatinid Phylogeny. American Museum Novitates, 3954: 1-29
New genus: Cretasquatina
New species: Cretasquatina americana
Abstract: Three-dimensional Late Cretaceous elasmobranch endoskeletal elements (including palatoquadrates, ceratohyals, braincase fragments, and a series of anterior vertebrae) are described from the Late Cretaceous University of Alabama Harrell Station Paleontological Site (HSPS), Dallas County, Alabama. The material is referred to the extant elasmobranch Family Squatinidae on the basis of several distinctive morphological features. It also exhibits features not shared by any modern or fossil Squatina species or the extinct Late Jurassic squatinid Pseudorhina. A new genus and species is erected, despite there being some uncertainty regarding potential synonymy with existing nominal species previously founded on isolated fossil teeth (curiously, no squatinid teeth have been documented from the HSPS). A preliminary phylogenetic analysis suggests that the new genus falls on the squatinid stem, phylogenetically closer to Squatina than Pseudorhina. The craniovertebral articulation in the new genus exhibits features considered convergent with modern batomorphs (skates and rays), including absence of contact between the posterior basicranium and first vertebral centrum, and a notochordal canal which fails to reach the parachordal basicranium. Supporting evidence that similarities in the craniovertebral articulation of squatinoids and batomorphs are convergent rather than synapomorphic (as "hypnosqualeans") is presented by an undescribed Early Jurassic batomorph, in which an occipital hemicentrum articulates with the first vertebral centrum as in all modern sharklike (selachimorph) elasmobranchs. The fossil suggests instead that the batomorph synarcual evolved by fusion of the anterior basiventral and basidorsal cartilages prior to the reduction of the anterior centra and loss of the occipital hemicentrum, not afterward as predicted by the hypnosqualean hypothesis.

LEBEDEV, O.A. & IVANOV, A.O. & LINKEVICH, V.V. (2020): Chondrichthyan spines from the Famennian (Upper Devonian) of Russia. Acta Geologica Polonica, 70: in press
New species: Sculptospina makhlaevi
Abstract: Very rare chondrichthyan spines from the Famennian (Upper Devonian) of European Russia are referred here to ctenacanthiforms, euselachians and a chondrichthyan group of uncertain systematic position. Ctenacanthus Agassiz, 1837 is recorded from the lower and middle Famennian of the central and north-western parts of the area. Sculptospina makhlaevi Lebedev gen. et sp. nov. originates from the lower Famennian of the Lipetsk Region. The holotype of ‘Ctenacanthus’ jaekeli Gross, 1933 and a new specimen from the upper Famennian of the South Urals are shown to belong to the same taxon, which is transferred to Acondylacanthus St. John and Worthen, 1875. New specimens of Tuberospina nataliae Lebedev, 1995 from the upper Famennian of Central Russia are described in detail. The newly presented material increases our knowledge of the composition of Famennian marine assemblages from the East European Platform. It is suggested that these assemblages may be classified as chondrichthyan-dominated and dipnoan-dominated. Hypothetically, after the end-Devonian Hangenberg extinction event, which affected numerous secondary consumers in vertebrate communities, some chondrichthyan groups could have encroached to take advantage of previously occupied ecological niches. Ctenacanthus, as well as Acondylacanthus and Amelacanthus survived the end-Devonian mass extinction to continue into the Carboniferous.

SAMBOU, B.S. & HAUTIER, L. & SARR, R. & TABUCE, R. & LIHOREAU, F. & THIAM, M. & LEBRUN, R. & MARTIN, J.E. & CAPPETTA, H. & ADNET, S. (2020): Contribution to the reappraisal of the mid Paleogene ichtyofauna of Western Africa with three new enigmatical elasmobranchs from Thanetian–Lutetian of Senegal. Annales de Paléontologie, in press
New genus: Atlantitrygon, Microtriftis
New species: Atlantitrygon senegalensis, Microtriftis matamensis, Odontorhytis priemi
Abstract: We report here three new elasmobranch fossil taxa from Thanetian–Lower Lutetian nearshore marine deposits of northeastern (Matam region) and central-western (Sine-Saloum region) Senegal. These three new taxa represent the oldest species of the enigmatical elasmobranch Odontorhytis, the oldest putative representatives of marine potamotrygonid, and an uncertain dasyatoid genus with the smallest grinding dentition ever described. These new taxa, representing the second Cenozoic elasmobranch remains formally described from Senegal, broaden our understanding of their evolutionary and biogeographic history in the equatorial Eastern Atlantic during this time period. Their occurrences confirm hypothesised stratigraphically correlations between the top of Matam Fm. and the base of Lam Lam Fm., refine the marine connections between these two Senegalese regions, and suggest that these genera were broadly distributed within the shallow marine settings of the equatorial Eastern Atlantic during latest Paleocene–early Middle Eocene.


WARREN, M.B. & BAKENHASTER, M.D. & SCHARER, R.M. & POULAKIS, G.R. & BULLARD, S.A. (2020): A new genus and species of fish blood fluke, Achorovermis testisinuosus gen. et sp. n. (Digenea: Aporocotylidae), infecting critically endangered smalltooth sawfish, Pristis pectinata (Rhinopristiformes: Pristidae) in the Gulf of Mexico. Folia Parasitologica, 67: 009
New genus: Achorovermis
New species: Achorovermis testisinuosus
AbstractAchorovermis testisinuosus gen. et sp. n. (Digenea: Aporocotylidae) infects the heart of the smalltooth sawfish, Pristis pectinata Latham (Rhinopristiformes: Pristidae), in the eastern Gulf of Mexico. Specimens of the new genus, along with the other blood flukes that infect batoids are similar by having an inverse U-shaped intestine and a curving testis as well as by lacking tegumental spines. The new genus differs from all of the other blood flukes infecting batoids by having an elongate body (>50 × longer than wide), a testis having >100 curves, and an ovary wholly anterior to the uterus. It differs from Ogawaia glaucostegi Cutmore, Cribb et Yong, 2018, the only other blood fluke infecting a rhinopristiform, by having a body that is >50 × (vs <30 ×) longer than wide, a testis that is >75 × (vs <40 ×) longer than wide and has >100 (vs <70) curves, an ovary wholly anterior to (vs lateral and dorsal to) the seminal vesicle, a uterus wholly posterior to (vs overlapping and lateral to both) the testis and ovary, and a sinuous (vs convoluted) uterus. The new species joins a small group of chondrichthyan blood flukes that lack tegumental spines: O. glaucostegiOrchispirium heterovitellatum Madhavi et Rao, 1970, Myliobaticola richardheardi Bullard et Jensen, 2008, Electrovermis zappum Warren et Bullard, 2019. Blood flukes infecting batoids are further unique by having a curving testis. That is, the blood flukes infecting species within Selachii are morphologically distinct from those infecting species within the Batoidea (excluding Gymnurahemecus bulbosus Warren et Bullard, 2019). Based on the morphological similarity, we suspect that the new species shares a recent common ancestor with O. glaucostegi. The discovery of the new species brings the total number of chondrichthyan blood flukes to 11 species assigned to nine genera.


PLEASE send your new papers to
juergen.pollerspoeck@shark-references.com or 

Latest Research Articles

Extant Chondrichthyes:
ARAYA, J.F. & REYES, P. & HUNE, M. (2020) On the presence of the Eastern Pacific Black Ghostshark Hydrolagus melanophasma (Chondrichthyes: Chimaeridae) in northern Chile, with notes on its distribution in the Eastern Pacific. Thalassas, in press https://dx.doi.org/10.1007/s41208-020-00220-0
ARNES-URGELLES, C. & PAEZ-ROSAS, D. & BARAHONA, D. & SALINAS-DE-LEON, P. (2020) First Direct Evidence of a Galapagos Sea Lion (Zalophus wollebaeki) Predated by a Galapagos Shark (Carcharhinus galapagensis). Aquatic Mammals, 46 (3): 254-258 https://dx.doi.org/10.1656/045.027.0207
BEGUM, A. & SARKER, S. & UZZAMAN, M.S. & SHAMSUZZAMAN, M.M. & ISLAM, M.M. (2020) Marine megafauna in the northern Bay of Bengal, Bangladesh: Status, threats and conservation needs. Ocean & Coastal Management, 192: 105228 https://dx.doi.org/10.1016/j.rsma.2020.101321
BERNARDO, C. & ADACHI, BAMCD, DA CRUZ, V.P. & FORESTI, F. & LOOSE, R.H. & BORNATOWSKI, H. (2020) The label "Cacao" is a shark or a ray and can be a threatened species! Elasmobranch trade in Southern Brazil unveiled by DNA barcoding. Marine Policy, 116: 103920 https://dx.doi.org/10.1016/j.ocecoaman.2020.105228
BOCKUS, A.B. & LABRECK, C.J. & CAMBERG, J.L. & COLLIE, J.S. & SEIBEL, B.A. (2020) Thermal Range and Physiological Tolerance Mechanisms in Two Shark Species from the Northwest Atlantic. Biological Bulletin, 238 (2): 131-144 https://dx.doi.org/10.1093/conphys/coaa041
CHAVEZ, E.J. & ARAUZ, R. & HEARN, A. & NALESSO, E. & STEINER, T. (2020) Asociación de tiburones con el Monte Submarino Las Gemelas y primera evidencia de conectividad con la Isla del Coco, Pacífico de Costa Rica [Association of sharks with Las Gemelas Seamount and first evidence of connectivity with Cocos Island, Pacific of Costa Rica]. Revista De Biologia Tropical, 68: 320-329
COFFEY, D.M. & ROYER, M.A. & MEYER, C.G. & HOLLAND, K.N. (2020) Diel patterns in swimming behavior of a vertically migrating deepwater shark, the bluntnose sixgill (Hexanchus griseus). Plos One, 15 (1): e0228253 https://dx.doi.org/10.1371/journal.pone.0228280
CURREY-RANDALL, L.M. & CAPPO, M. & SIMPFENDORFER, C.A. & FARABAUGH, N.F. & HEUPEL, M.R. (2020) Optimal soak times for Baited Remote Underwater Video Station surveys of reef-associated elasmobranchs. Plos One, 15 (5): e0231688 https://dx.doi.org/10.14411/fp.2020.009
CURRIE, J.C. & ATKINSON, L.J. & SINK, K.J. & ATTWOOD, C.G. (2020) Long-Term Change of Demersal Fish Assemblages on the Inshore Agulhas Bank Between 1904 and 2015. Frontiers in Marine Science, 7: 355 https://dx.doi.org/10.3389/fmars.2020.00422
DA SILVA, J.P.C.B. & DATOVO, A. (2020) The coracoid bar and its phylogenetic importance for elasmobranchs (Chondrichthyes). Zoologischer Anzeiger, in press  https://dx.doi.org/10.1016/j.jsames.2020.102566
DOLTON, H.R. & GELL, F.R. & HALL, J. & HALL, G. & HAWKES, L.A. & WITT, M.J. (2020) Assessing the importance of Isle of Man waters for the basking shark Cetorhinus maximus. Endangered Species Research, 41: 209-223 https://dx.doi.org/10.3389/fmars.2020.00291
DUNN, N. & JOHRI, S. & CURNICK, D. & CARBONE, C. & DINSDALE, E.A. & CHAPPLE, T.K. & BLOCK, B.A. & SAVOLAINEN, V. (2020) Complete mitochondrial genome of the gray reef shark, Carcharhinus amblyrhynchos (Carcharhiniformes: Carcharhinidae). Mitochondrial DNA Part B, 5 (3): 2080-2082 https://dx.doi.org/10.1080/23802359.2020.1765209
ELSTON, C. & COWLEY, P.D. & VON BRANDIS, R.G. & FISK, A. (2020) Dietary niche differentiation in a mesopredatory dasyatid assemblage. Marine Biology, 167: 89 https://dx.doi.org/10.1007/s10641-020-00976-7
GRAINGER, R. & PEDDEMORS, V.M. & RAUBENHEIMER, D. & MACHOVSKY-CAPUSKA, G.E. (2020) Diet Composition and Nutritional Niche Breadth Variability in Juvenile White Sharks (Carcharodon carcharias). Frontiers in Marine Science, 7: 422 https://dx.doi.org/10.3390/su12072683
HARDIMAN, N. & BURGIN, S. & SHAO, J. (2020) How Sharks and Shark-Human Interactions are Reported in Major Australian Newspapers. Sustainability, 12 (7): 2683 https://dx.doi.org/10.3856/vol48-issue1-fulltext-2176
HARRIS, J.L. & MCGREGOR, P.K. & OATES, Y. & STEVENS, G.M.W. (2020) Gone with the wind: Seasonal distribution and habitat use by the reef manta ray (Mobula alfredi) in the Maldives, implications for conservation. Aquatic Conservation-Marine and Freshwater Ecosystems, in press https://dx.doi.org/10.1002/jmor.21123
HOGLIN, B.E. & MINER, M. & DORES, R.M. (2020) Pharmacological properties of whale shark (Rhincodon typus) melanocortin-2 receptor and melancortin-5 receptor: Interaction with MRAP1 and MRAP2. General and Comparative Endocrinology, 293: 113463 https://dx.doi.org/10.1017/s0025315420000259
JOHRI, S. & DUNN, N. & CHAPPLE, T.K. & CURNICK, D. & SAVOLAINEN, V. & DINSDALE, E.A. & BLOCK, B.A. (2020) Mitochondrial genome of the Silvertip shark, Carcharhinus albimarginatus, from the British Indian Ocean Territory. Mitochondrial DNA Part B, 5 (3): 2085-2086 https://dx.doi.org/10.1086/708718
JOHRI, S. & TIWARI, A. & KERR, E.N. & DINSDALE, E.A. (2020) Mitochondrial genome of the Smoothnose wedgefish Rhynchobatus laevis from the Western Indian Ocean. Mitochondrial DNA Part B, 5 (3): 2083-2084 https://dx.doi.org/10.1080/23802359.2020.1765210
JONES, C.M. & DRIGGERS, W.B. & HANNAN, K.M. & HOFFMAYER, E.R. & JONES, L.M. & RAREDON, S.J. (2020) An annotated checklist of the chondrichthyan fishes inhabiting the northern Gulf
of Mexico Part 1: Batoidea. Zootaxa, 4803 (2): 281–315 https://dx.doi.org/10.1206/3954.1
KABASAKAL, H. (2020) A Field Guide to the Sharks of Turkish Waters. Turkish Marine Research Foundation (TUDAV) Publication No: 55, Istanbul, Turkey. 133 p.
KANAGUSUKU, K. & ROMERO, M. & RAMIREZ-AMARO, S. (2020) Multiple morphological abnormalities in a blue shark Prionace glauca (Linnaeus, 1758) embryo from the Peruvian coast, southeast Pacific. Latin American Journal of Aquatic Research, 48 (1): 141-145 https://dx.doi.org/10.3856/vol48-issue3-fulltext-2433
KROETZ, A.M. & MATHERS, A.N. & CARLSON, J.K. (2020) Evaluating protected species bycatch in the US Southeast Gillnet Fishery. Fisheries Research, 228: 105573 https://dx.doi.org/10.1016/j.fishres.2020.105577
KUMAR, A. & ADHAVAN, D. & VINUGANESH, A. & PRAKASH, S. (2020) DNA barcoding revealed first record of the 'fine spotted whipray' Himantura tutul (Myliobatoidei: Dasyatidae) in the Indian coastal waters. Journal of Applied Ichthyology, in press https://dx.doi.org/10.1111/joa.13212
LE CROIZIER, G. & LORRAIN, A. & SCHAAL, G. & KETCHUM, J. & HOYOS-PADILLA, M. & BESNARD, L. & MUNARON, J.M. & LE LOC'H, F. & POINT, D. (2020) Trophic resources and mercury exposure of two silvertip shark populations in the Northeast Pacific Ocean. Chemosphere, 253: 126645 https://dx.doi.org/10.1016/j.envpol.2020.114923
LOPEZ-GARRO, A. & ZANELLA, I. & GOLFIN-DUARTE, G. & PEREZ-MONTERO, M. (2020) Residencia del tiburón punta blanca de arrecife, Triaenodon obesus, (Carcharhiniformes: Carcharhinidae) en las bahías Chatham y Wafer del Parque Nacional Isla del Coco, Costa Rica [Residency of the whitetip reef shark (Triaenodon obelus) in Chatham and Wafer Bays, Isla del Coco National Park, Costa Rica].  Revista De Biologia Tropical, 68: 330-339
MABRAGANA, E. & GONZALEZ-CASTRO, M. & GABBANELLI, V. & VAZQUEZ, D.M. & DE ASTARLOA, J.M.D. (2020) Polymorphism in Conservative Structures? The Scapulocoracoids in Skates Genus Psammobatis (Chondrichthyes, Arhynchobatidae) and the Validity of P. parvacauda. Frontiers in Marine Science, 7: 291 https://dx.doi.org/10.3389/fmars.2020.00292
MARCONI, A. & HANCOCK-RONEMUS, A. & GILLIS, J.A. (2020) Adult chondrogenesis and spontaneous cartilage repair in the skate, Leucoraja erinacea. Elife, 9: e53414 https://dx.doi.org/
MARTINS, A.P.B. & HEUPEL, M.R. & BIERWAGEN, S.L. & CHIN, A. & SIMPFENDORFER, C. (2020) Diurnal activity patterns and habitat use of juvenile Pastinachus ater in a coral reef flat environment. Plos One, 15 (2): e0228280 https://dx.doi.org/10.1371/journal.pone.0230308
MARTINSA, M.F. & COSTA, P.G. & BIANCHINI, A. (2020) Contaminant screening and tissue distribution in the critically endangered Brazilian guitarfish Pseudobatos horkelii. Environmental Pollution, 265 B: 114923 https://dx.doi.org/10.1016/j.fishres.2020.105569
MEESE, E.N. & LOWE, C.G. (2020) Environmental effects on daytime sheltering behaviors of California horn sharks (Heterodontus francisci). Environmental Biology of Fishes, in press https://dx.doi.org/10.1007/s10750-020-04283-1
MENDOZA-PORTILLO, F.J. & RAMIREZ-RODRIGUEZ, M. & VARGAS-LOPEZ, V. (2020) Interactions of small-scale fisheries in Mexico's northwest Pacific. Latin American Journal of Aquatic Research, 48 (1): 94-105 https://dx.doi.org/10.3856/vol48-issue1-fulltext-2317
MENON, M. & MAHESWARUDU, G. & RAMULU, K.S. & KIZHAKUDAN, S.J. (2020) Reproductive biology and diet of the longtail butterfly ray Gymnura poecilura (Shaw, 1804) along western Bay of Bengal. Journal of the Marine Biological Association of the United Kingdom, 100 (3): 461-470 https://dx.doi.org/10.1038/s41598-020-66876-z
MICARELLI, P. & DI GRUMO, D. & REINERO, F. & GIGLIO, G. & SPERONE, E (2020) Observations about Not Invasive Method for Individual Identificatıon of Small Spotted Catshark (Scyliorhinus canicula, Linnaeus 1758) in Controlled Conditions. International Journal of Oceanography & Aquaculture, 4 (1): 000189 https://dx.doi.org/10.24425/agp.2020.132255
MOHAN, J.A. & JONES, E.R. & HENDON, J.M. & FALTERMAN, B. & BOSWELL, K.M. & HOFFMAYER, E.R. & WELLS, R.J.D. (2020) Capture stress and post-release mortality of blacktip sharks in recreational charter fisheries of the Gulf of Mexico. Conservation Physiology, 8: coaa041 https://dx.doi.org/10.1111/jai.14060
MOYER, J.K. & DODD, J. & IRSCHICK, D.J. (2020) Observation of a Sea Lamprey, Petromyzon marinus, on a Pelagic Blue Shark, Prionace glauca. Northeastern Naturalist, 27 (2): N16-N20 https://dx.doi.org/10.22201/ib.20078706e.2020.91.3054
MOYER, J.K. & FINUCCI, B. & RICCIO, M.L. & IRSCHICK, D.J. (2020) Dental morphology and microstructure of the Prickly Dogfish Oxynotus bruniensis (Squaliformes: Oxynotidae). Journal of Anatomy, in press https://dx.doi.org/10.1127/njgpa/2020/0908
MULL, C.G. & YOPAK, K.E. & DULVY, N.K. (2020) Maternal Investment, Ecological Lifestyle, and Brain Evolution in Sharks and Rays. American Naturalist, 195 (6): 1056-1069 https://dx.doi.org/10.7554/eLife.53414
NOBLE, R. (2020) Every Picture Tells a Story Using Data Visualization to Protect Great White Shark Population. Sea Technology, 61 (5): 14-16
PAGLIARINI, C.D. & RIBEIRO, C.D. & SPADA, L. & DELARIVA, R.L. & CHAGAS, J.M.A. & DOS ANJOS, L.A. & RAMOS, I.P. (2020) Trophic ecology and metabolism of two species of nonnative freshwater stingray (Chondrichthyes: Potamotrygonidae). Hydrobiologia, 847: 2895–2908 https://dx.doi.org/10.1007/s11160-020-09605-z
PAPASTAMATIOU, Y.P. & BRITTON, C. & BURGESS, G.H. (2020) Using side-scan sonar to survey critically endangered smalltooth sawfish. Fisheries Research, 228: 105577 https://dx.doi.org/10.1016/j.isci.2020.101205
PARMENTIER, E. & BANSE, M. & BOISTEL, R. & COMPERE, P. & BERTUCCI, F. & COLLEYE, O. (2020) The development of hearing abilities in the shark Scyliorhinus canicula. Journal of Anatomy, in press https://dx.doi.org/10.1111/joa.13251 
PETERSON, C.T. & GRUBBS, R.D. & MICKLE, A. (2020) Trophic ecology of elasmobranch and teleost fishes in a large subtropical seagrass ecosystem (Florida Big Bend) determined by stable isotope analysis. Environmental Biology of Fishes, in press https://dx.doi.org/10.1007/s10641-020-00977-6
RODRÍGUEZ-CABELLO, C. & PÉREZ, M. & GRASA, I. (2020) Taxonomic research on Deania calcea and Deania profundorum (Family: Centrophoridae) in the Cantabrian Sea (Northeast Atlantic) with comments on Deania hystricosa. Regional Studies in Marine Science, 37: 101321 https://dx.doi.org/10.1016/j.ygcen.2020.113463
SCHOEMAN, R.P. & PATTERSON-ABROLAT, C. & PLÖN, S. (2020) A Global Review of Vessel Collisions With Marine Animals. Frontiers in Marine Science, 7: 292 https://dx.doi.org/10.3389/fmars.2020.00355
SEN, S. & DASH, G. & KIZHAKUDAN, S.J. & CHAKRABORTY, R.D. & MUKHERJEE, I. (2020) New record of the giant freshwater whipray, Urogymnus polylepis from West Bengal waters, east coast of India. Ichthyological Exploration of Freshwaters, 30 (1): 91-95 https://dx.doi.org/10.23880/ijoac-16000189
SEN, S. & DASH, G. & VALAPPIL, A.K. & KIZHAKUDAN, S.J. & CHAKRABORTY, R. (2020) Occurrences of Intersexual Hound Sharks, Iago cf. omanensis (Triakidae: Carcharhiniformes) from North-western Bay of Bengal. Thalassas, in press https://dx.doi.org/10.1016/j.biocon.2020.108459
SHEA, B.D. & BENSON, C.W. & DE SILVA, C. & DONOVAN, D. & ROMEIRO, J. & BOND, M.E. & CREEL, S. & GALLAGHER, A.J. (2020) Effects of exposure to large sharks on the abundance and behavior of mobile prey fishes along a temperate coastal gradient. Plos One, 15 (3): e0230308 https://dx.doi.org/10.1371/journal.pone.0231688
SHIFFMAN, D.S. & BITTICK, S.J. & CASHION, M.S. & COLLA, S.R. & CORISTINE, L.E. & DERRICK, D.H. & GOW, E.A. & MACDONALD, C.C. & O’FERRALL, M.M. & OROBKO, M. & POLLOM, R.A. & PROVENCHER, J. & DULVY, N.K. (2020) Inaccurate and Biased Global Media Coverage Underlies Public Misunderstanding of Shark Conservation Threats and Solutions. iScience, in press https://dx.doi.org/10.1016/j.jcz.2020.06.004
SIMPSON, S.J. & HUMPHRIES, N.E. & SIMS, D.W. (2020) The spatial ecology of Rajidae from mark-recapture tagging and its implications for assessing fishery interactions and efficacy of Marine Protected Areas. Fisheries Research, 228: 105569 https://dx.doi.org/10.1016/j.fishres.2020.105573
SNODGRASS, D.J.G. & ORBESEN, E.S. & WALTER, J.F. & HOOLIHAN, J.P. & BROWN, C.A. (2020) Potential impacts of oil production platforms and their function as fish aggregating devices on the biology of highly migratory fish species. Reviews in Fish Biology and Fisheries, in press https://dx.doi.org/10.1007/s12237-019-00600-7
SOARES, K.D.A. (2020) Comparative anatomy of the clasper of catsharks and its phylogenetic implications (Chondrichthyes: Carcharhiniformes: Scyliorhinidae). Journal of Morphology, 281 (6): 591-607 https://dx.doi.org/10.1007/s00227-020-03695-w
SOARES, K.D.A. & DE CARVALHO, M.R. (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution, 96 (2): 345-395 https://dx.doi.org/10.5061/dryad.kprr4xh1b
SPAET, J.L.Y. & PATTERSON, T.A. & BRADFORD, R.W. & BUTCHER, P.A. (2020) Spatiotemporal distribution patterns of immature Australasian white sharks (Carcharodon carcharias). Scientific Reports, 10: 10169 https://dx.doi.org/10.1071/is19048
STRICKLAND, B.A. & MASSIE, J.A. & VIADERO, N. & SANTOS, R. & GASTRICH, K.R. & PAZ, V. & O'DONNELL, P. & KROETZ, A.M. & HO, D.V.T. & REHAGE, J.S. & HEITHAUS, M.R. (2020) Movements of Juvenile Bull Sharks in Response to a Major Hurricane Within a Tropical Estuarine Nursery Area. Estuaries and Coasts, 43 (5): 1144-1157 https://dx.doi.org/10.1007/s12542-020-00518-7
TORRES-HUERTA, A.M. & CRUZ-ACEVEDO, E. & CARRASCO-BAUTISTA, P.E. & MERAZ-HERNANDO, J.F. & RAMÍREZ-CHÁVEZ, E.J. & TAPIA-GARCÍA, M. & GARCÍA, A. (2020) Reproductive ecology of the witch guitarfish Zapteryx xyster Jordan & Evermann, 1896 (Chondrichtyes: Trygonorrhinidae) in the Gulf of Tehuantepec, Mexican Pacific. Marine and Freshwater Research, 71 (7): 844-854 https://dx.doi.org/10.1080/23802359.2020.1765208
VIANA, S.T.F.L. & DE CARVALHO, M.R. (2020) Squalus shiraii sp. nov. (Squaliformes, Squalidae), a new species of dogfish shark from Japan with regional nominal species revisited. Zoosystematics and Evolution, 96 (2): 275-311 https://dx.doi.org/10.3897/zse.96.52420
VIANA-MORAYTA, J.E. & TORRES-ROJAS, Y.E. & CAMALICH-CARPIZO, J. (2020) Diet shifts of Rhizoprionodon terraenovae from the southern Gulf of Mexico: possible scenario by temperature changes. Latin American Journal of Aquatic Research, 48 (3): in press https://dx.doi.org/10.3897/zse.96.51962
WALLS, R.H.L. & DULVY, N.K. (2020) Eliminating the dark matter of data deficiency by predicting the conservation status of Northeast Atlantic and Mediterranean Sea sharks and rays. Biological Conservation, 246: Unsp 108459 https://dx.doi.org/10.1016/j.chemosphere.2020.126645
WHITEHEAD, D.A. & JAKES-COTA, U. & PANCALDI, F. & GALVAN-MAGANA, F. & GONZALEZ-ARMAS, R. (2020) The influence of zooplankton communities on the feeding behavior of whale shark in Bahia de La Paz, Gulf of California. Revista Mexicana De Biodiversidad, 91 (1): UNSP e913054  https://dx.doi.org/10.2298/GABP200214003T
Extinct Chondrichthyes:

DUFFIN, C. & LARS, J. (2020) A fin spine of Recurvacanthus (Myriacanthidae, Holocephali) from the Posidonienschiefer (Early Jurassic) of SW Germany. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen Band, 296 (3): 317-326  https://dx.doi.org/10.11646/zootaxa.4803.2.3
EVERAERT, S. (2020) Parotodus benedenii (Le Hon, 1871) uit het Zand van Antwerpen (Midden Mioceen) te Berchem (Antwerpen, België). Afzettingen WTKG, 41 (2): 36-40
LEBEDEV, O.A. & IVANOV, A.O. & LINKEVICH, V.V. (2020) Chondrichthyan spines from the Famennian (Upper Devonian) of Russia. Acta Geologica Polonica, 70: in press https://dx.doi.org/10.3354/dao03458
LOPES, R.P. & PEREIRA, J.C. & DILLENBURG, S.R. & TATUMI, S.H. & YEE, M. & FIGUEIREDO, A.M.G. & KINOSHITA, A. & BAFFA, O. (2020) Late Pleistocene-Holocene fossils from Mirim Lake, Southern Brazil, and their paleoenvironmental significance: I - Vertebrates. Journal of South American Earth Sciences, 100: 102566  https://dx.doi.org/10.1016/j.marpol.2020.103920
MAISEY, J.G. & EHRET, D.J. & DENTON, J.S.S. (2020) A New Genus of Late Cretaceous Angel Shark (Elasmobranchii; Squatinidae), with Comments on Squatinid Phylogeny. American Museum Novitates, 3954: 1-29 https://dx.doi.org/10.1371/journal.pone.0228253
SAMBOU, B.S. & HAUTIER, L. & SARR, R. & TABUCE, R. & LIHOREAU, F. & THIAM, M. & LEBRUN, R. & MARTIN, J.E. & CAPPETTA, H. & ADNET, S. (2020) Contribution to the reappraisal of the mid Paleogene ichtyofauna of Western Africa with three new enigmatical elasmobranchs from Thanetian–Lutetian of Senegal [Contribution à la réévaluation de l’ichtyofaune du Paléogène moyen en Afrique de l’Ouest avec trois nouveaux élasmobranches énigmatiques du Thanétien–Lutétien du Sénégal]. Annales de Paléontologie, in press
TAKAKUWA, Y. & HIROSE, K. & KUROSU, H. (2020) The first occurrence of fossil sharpnose sevengill shark (Heptranchias; Hexanchiformes) from the Eocene Sakasegawa Formation, Amakusa, Kumamoto, Japan. Bulletin of Gunma Museum of Natural History, 24: 25-30
TRIF, N. & GHAEMI, F. & TAHERI, J. & TAHERPOUR-KHALIL-ABAD, M. (2020) Short note on the first record of fossil shark teeth in the Chehel-Kaman formation, Iran. Geoloski anali Balkanskoga poluostrva, in press  https://dx.doi.org/10.23788/ief-1113
VILLAFAÑA, J.A. & MARRAMÀ, G. & KLUG, S. & POLLERSPÖCK, J. & BALSBERGER, M. & RIVADENEIRA, M. & KRIWET, J. (2020) Sharks, rays and skates (Chondrichthyes, Elasmobranchii) from the Upper Marine Molasse (middle Burdigalian, early Miocene) of the Simssee area (Bavaria, Germany), with comments on palaeogeographic and ecological patterns. Paläontologische Zeitschrift, in press  https://dx.doi.org/10.1007/s41208-020-00209-9


BURRESON, E.M. (2020) Marine and estuarine leeches (Hirudinida : Ozobranchidae and Piscicolidae) of Australia and New Zealand with a key to the species.-Invertebrate Systematics, 34 (3): 235-259 www.dx.doi.org/10.1071/is19048
MARTIN, S.B. & DOWNIE, A.J. & CRIBB, T.H. (2020) A new subfamily for a clade of opecoelids (Trematoda: Digenea) exploiting marine fishes as second-intermediate hosts, with the first report of opecoelid metacercariae from an elasmobranch.-Zoological Journal of the Linnean Society, 188 (2): 455-472
MORAVEC, F. & DALRYMPLE, K.M. & GALLOWAY, A.S. & BARKER, A.M. & DE BURON, I. (2020) First record of Piscicapillaria bursata (Nematoda: Capillariidae), a parasite of hammerhead sharks Sphyrna spp., in the western Atlantic Ocean.-Diseases of Aquatic Organisms, 138: 133-136 www.dx.doi.org/10.3354/dao03458
WARREN, M.B. & BAKENHASTER, M.D. & SCHARER, R.M. & POULAKIS, G.R. & BULLARD, S.A. (2020) A new genus and species of fish blood fluke, Achorovermis testisinuosus gen. et sp. n. (Digenea: Aporocotylidae), infecting critically endangered smalltooth sawfish, Pristis pectinata (Rhinopristiformes: Pristidae) in the Gulf of Mexico.-Folia Parasitologica, 67: 009 www.dx.doi.org/10.14411/fp.2020.009



A new character for Pokémon? Novel endemic dogfish shark species discovered from Japan

Date: June 11, 2020
Source: Pensoft Publishers
Summary: A new endemic deep-water dogfish shark: Squalus shiraii, was discovered in the tropical waters of Southern Japan by an international team of scientists. The finding brings the amount of spurdogs shark species inhabiting Japanese waters to six.

Great white shark diet surprises scientists

Understanding how sharks feed is vital for managing human interactions

Date: June 7, 2020
Source: University of Sydney
Summary: The first-ever detailed analysis of the diet of great white sharks has shown they spend more time feeding at the seafloor than many would have expected.