NEWSLETTER 06/2020 03.06.2020

Please acknowledge use of the database www.shark-references.com in your publications, and cite: 

Pollerspöck, J. & Straube, N. 2020, Bibliography database of living/fossil sharks, rays and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali), www.shark-references.com, World Wide Web electronic publication, Version 2020

New project at shark-references!

for more information check the pdf or video

and a new paper (co-authored by Jürgen team shark-references) is just out

free download via 


Name: Arturo Angulo Sibaja
Affiliation: Colección Ictiológica, Museo de Zoología, Universidad de Costa Rica
Your interests in Chondrichthyans or short introduction to your research (max. 90 words): My research interests center around systematics, taxonomy, biogeography and ecology of sharks, rays and chimaeras from the Tropical Eastern Pacific. In the last years, in collaboration with some colleagues, I have described some new species within such groups and updated the national (Costa Rica) and regional lists.

Would you like to become a shark-reference partner? Please contanct us per E-mail!

Partner in Google-Maps




New Images

Many thanks to the following people for providing images:

Frederik Mollen (Elasmobranch Research Belgium) for the images of Dipturus oxyrinchus (LINNAEUS, 1758)

G. M. MASUM BILLAH, Program Coordinator- Marine Mega-Fauna & Fisheries, Wildlife Conservation Society- Bangladesh Program (masumwcsbd@gmail.com) for images of Hemipristis elongata (KLUNZINGER, 1871) and Gymnura poecilura (SHAW, 1804)

Bryan Huerta for a image of Mustelus sinusmexicanus HEEMSTRA, 1997

Pradip Patade, India for images of Glaucostegus granulatus (CUVIER, 1829)Carcharhinus limbatus (MÜLLER & HENLE, 1839) and Rhizoprionodon oligolinx SPRINGER, 1964

Charlie Underwood, Natural History Museum, London for images of e.g. Sclerorhynchus atavus WOODWARD, 1889

Tiffany Slater for a image of the holotype of Paracestracion danieli nov. sp. (PBP-SOL-0005)

Gerrica Hightower, Australia for a image of Pastinachus ater (MACLEAY, 1883)

Missing papers:

Many thanks to all friends of shark-references, who sent us some missing papers last month!

Shark-References would kindly like to ask you for your contribution to this project.

At the moment we are looking for some of the following papers:

Extinct Chondrichthyes:
NEWTON, E.T.  (1878) The chimeroid fishes of the British Cretaceous rocks. Memoirs of the Geological Survey of the United Kingdom, Monograph, 4: 1–64

BONARELLI, G.  (1898) I fossili senoniani dell'Appennino centrale che si conservano a Peongia nella collezione Bellucci. Atti della Reale Accademia delle scienze di Torino, 34: 1020–1027, pl.

WEILER, W.  (1930) Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wüsten Ägyptens. VI. Beschreibung von Wirbeltier-Resten aus dem nubischen Sandsteine Oberägyptens und aus ägyptischen Phosphaten nebst Bemerkungen über die Geologie der Umgegend von Mahamid in Oberägypten.  Abhandlungen der Königlich Bayerischen Akademie der Wissenschaften, Abt., N.F., 7: 1–42, 4 pl.

CHECCHIA-RISPOLI, B. (1933) Di un nuevo genere di «Pristidae» del Cretaceo superiore della Tripolitania.    Memorie della Reale Accademia italiana, Classe Scienze Fisiche Mathematiche e Naturale, 4 (1): 1–6, 1 pl.

MATSUMOTO, H. (1936) Upper Miocene vertebrates from Kumanodô, Natori district, province of Rikuzen. Dobutsugaku Zasshi, 48: 475–480, 5 fig.

FOWLER, H.W. (1947) New taxonomic names of fish-like vertebrates. Notulae Naturae, 187: 1–16

ALVINERIE, J. & ANDREIEFF, P. & ANGLADA, R. & AUBERT, J. & CAPPETTA, H. & CARALP, M. & CARATINI, C. & CARBONNEL, G. & CATZIGRAS, F. & COURME-RAULT, M.-D. & CHATEAUNEUF, J.-J. & DEMARCQ, G. & DUCASSE, O. & FATTON, E. & GLAÇON, G. & LABRACHERIE, M. & LAURIAT, A. & LE CALVEZ, Y. & LORENZ, C. & MAGNE, J. & MARGEREL, J.-P. & POIGNANT, A. & PUJOL, C. & ROGER, J. & ROMAN, J. & BLONDEAU, A. & MULLER, C. (1973) A propos de la limite oligo-miocène: résultats préliminaires d'une recherche collective sur les gisements d'Escornébéou (Saint-Géours-de-Maremne, Landes, Aquitaine méridionale). Présence de Globigerinoides dans les faunes de l'Oligocène supérieur. Comptes rendus sommaires des séances de la Société géologique de France: 75–76

BOYD, B.M. (2016) Fossil sharks and rays of Gainesville creeks; Alachua County, Florida: Hogtown group; (middle Miocene to lower Pliocene). Florida Paleontological Society, Special Publication

Extant Chondrichthyes:

KAMOHARA, T. (1943) Some unrecorded and two new fishes from Prov. Tosa, Japan. Bulletin of the Biogeographical Society of Japan, 13 (17): 125–137

DE BUEN, F. (1950) Contribuciones a la Ictiología. II. El tiburón vitamínico de la costa uruguaya Galeorhinus vitaminicus nov. sp., y algunas consideraciones generales sobre su biología. Publicaciones Cientificas, Servicio Oceanografico y de Pesca, Ministerio de Industrias y Trabajo, Montevideo No. 4: 153–162.

CADENAT, J. (1951) Initiations Africaines. III. Poissons de Mer du Sénégal. Institute Francais d'Afrique Noire. Initiations Africaines. III. Poissons de Mer du Sénégal.: 1–345

WEIBEZAHN, F.H. (1953) Una nueva especie de Scyliorhinus de Venezuela (Chondrichthyes - Elasmobranchii). Novedades cientificas. Serie zoológica. Museo de Historia Natural La Salle, 9: 1–7.

SMITH, J.L.B. (1958) The mystery killer, the new shark Carcharhinus vanrooyeni. Veld & Vlei, 3 (9): 12–14, 28.

SICCARDI, E. (1961) Cetorhinus en el Atlantico sur (Elasmobranchii: Cetorhinidae). Actas y trabajos del Primer Congreso Sudamericano de Zoologia, 4 (5): 251–263

GUBANOV, E.P. & SCHLEIB, N.A. (1980) Sharks of the Arabian Gulf. Kuwait Ministry of Public Works, Agracultural Department, Fisheries Division. Sharks of the Arabian Gulf.: 1–69

DOLGANOV, V.N. (1983) Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod. [Manual for identification of cartilaginous fishes of Far East seas of USSR and adjacent waters.] TINRO, Vladivostok. Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod.: 92 pp.

Please support www.shark-references.com and send missing papers (not listed papers or papers without the info-symbol) to juergen.pollerspoeck@shark-references.com or nicolas.straube@shark-references.com



Upcoming Meetings:

The annual scientific conference of the European Elasmobranch Association 2020 will be held in Leiden, the Netherlands between 28th and 30th October, 2020. Venue is the fascinating Naturalis Biodiversity Center.



IMCC6 to be held online

In light of the challenges the global community faces from the covid-19 pandemic, we have decided change from an in-person meeting to an online IMCC6.


The 6th International Marine Conservation Congress

To conserve the world’s oceans we must go beyond science and use it to inform policy and management to catalyze change. The International Marine Conservation Congress (IMCC) brings together conservation professionals and students to develop new and powerful tools to further marine conservation science and policy.

With over 700 marine conservation professionals and students in attendance, IMCC is the most important international event for anyone involved or interested in marine conservation. The meeting brings together marine conservationists from many walks of life including but not limited to scientists, practitioners, educators, policy-makers, artists and journalists.

IMCC6 takes place in the sailing city of Kiel, Germany. Join us in Kiel as we come together to help Make Marine Science Matter!

for more information please visit https://conbio.org/mini-sites/imcc6



A space for the dissemination of knowledge and interaction among researchers interested in sharks, rays and chimeras, both in Colombia and in Latin America. It is done every two years and since 2008 six versions have been made.

It will be held at the Pontificia Universidad Javeriana de Cali, Colombia,
from July 6 to 10, 2020.



Eighth International Meeting on the enhancement and preservation of paleontological heritage, at the science faculty of El Jadida, Morocco from November 25 to 27, 2020 (visit the web page : http://www.fs.ucd.ac.ma/fs/riv3p8).



Extant Chondrichthyes:

no taxonomic news this month

Extinct Chondrichthyes:

 (2020): Evolutionary relationships among bullhead sharks (Chondrichthyes, Heterodontiformes). Papers in Palaeontology, in press
New species: Paracestracion danieli
Abstract: The evolution of modern sharks, skates and rays (Elasmobranchii) is largely enigmatic due to their possession of a labile cartilaginous skeleton; consequently, taxonomic assignment often depends on isolated teeth. Bullhead sharks (Heterodontiformes) are a group of basal neoselachians, thus their remains and relationships are integral to understanding elasmobranch evolution. Herein we fully describe †Paracestracion danieli – a bullhead shark from the Late Jurassic plattenkalks of Eichstätt, Germany (150–154 Ma) – for its inclusion in cladistic analysis (utilizing parsimonious principles) of morphological characters from complete †Paracestracion and Heterodontus fossil specimens as well as extant forms of the latter. The presence of two separate monophyletic clades within Heterodontiformes was confirmed, based on predominantly non‐dental characters, which show a strong divergence in body morphology between †Paracestracion and Heterodontus ; the latter possessing a first dorsal fin and pectoral fins that are more anterior and pelvic fins that are more posterior. This study emphasizes the importance of including non‐dental features in heterodontiform systematics (as compared with the use of dental characters alone) and supports the erection of the family †Paracestracionidae. Further, phylogenetic analysis of molecular data from five extant species suggests that crown heterodontiforms arose from a diversification event 42.58 Ma off the west coast of the Americas.


SHAMSI, S. & BARTON, D.P. & ZHU, X.C. (2020): Description and genetic characterisation of Pulchrascaris australis n. sp. in the scalloped hammerhead shark, Sphyrna lewini (Griffin & Smith) in Australian waters. Parasitology Research, in press
New species: Pulchrascaris australis
Abstract: Being listed in the IUCN Red List of Threatened Species, knowledge on the biology, health and diseases of the scalloped hammerhead, Sphyrna lewini (Griffith & Smith) is limited; this is especially true for its parasites. In this paper, a new species, Pulchrascaris australis, is morphologically described followed by genetic characterisation based on the sequence of the ITS region. The new species can be easily differentiated from its congeners based on the morphology of the mouthpart, spicules, plectanes, eggs and vulva. Phylogenetic analyses clearly distinguish specimens in the present study from other parasitic nematodes found in the Australasian waters and elsewhere. The sequencing data also suggest that Terranova larval type I found previously in various fish from New Caledonian and Australian (Queensland) waters also belong to Pulchrascaris australis n. sp.
BRAY, R.A. & WAESCHENBACH, A. & LITTLEWOOD, D.T.J. & HALVORSEN, O. & OLSON, P.D. (2020): Molecular circumscription of new species of Gyrocotyle Diesing, 1850 (Cestoda) from deep-sea chimaeriform holocephalans in the North Atlantic. Systematic Parasitology, 97 (3): 285-296
New species: Gyrocotyle haffii, Gyrocotyle discoveryi
Abstract: Chimaeras, or ratfishes, are the only extant group of holocephalan fishes and are the sole host group of gyrocotylidean cestodes, which represent a sister group of the true tapeworms (Eucestoda). These unique, non-segmented cestodes have been known since the 1850s and multiple species and genera have been erected despite a general agreement that the delineation of species on the basis of morphology is effectively impossible. Thus, in the absence of molecular studies, the validity of gyrocotylid taxa and their specific host associations has remained highly speculative. Here we report the presence of Gyrocotyle spp. from rarely-caught deep-sea chimaeras collected in the North-East Atlantic, and describe two new species: G. haffii n. sp. from the bent-nose chimaera, Harriota raleighana Goode & Bean, and G. discoveryi n. sp. from the large-eyed rabbit fish, Hydrolagus mirabilis (Collett). Nuclear ribosomal sequence data were generated for individual parasites taken from different host species collected on different dates and from different localities and were combined with previously published sequences. Phylogenetic analyses supported the recognition of independent lineages and clusters, indicative of species, but were indecisive in recovering the root of the tree in analyses that included non-gyrocotylid outgroup taxa. The molecular data reveal variation not reflected in morphology and point to a complex picture of genetic divergence shaped by both isolation and migration in the deep-sea environment.
ZARAGOZA-TAPIA, F. & PULIDO-FLORES, G. & MONKS, S. (2020): Three new species of Acanthobothrium Blanchard, 1848 (Cestoda: Onchoproteocephalidea) in Stingrays (Dasyatidae) from the Pacific coast in Mexico. Zootaxa, 4766 (1): 157-172
New species: Acanthobothrium ppdeleoni, Acanthobothrium hypanus, Acanthobothrium sinaloaensis
Abstract: Three new species of Acanthobothrium Blanchard, 1848 (Onchoproteocephalidea: Onchobothriidae) are described from Pacific coastal waters of Mexico. Based on four criteria for categorization of species of Acanthobothrium, the three new species belong to Category 2 species, characterized by their total length (< 15 mm), number of proglottids (< 50) and testes (<80), and with asymmetrical ovary. Acanthobothrium ppdeleoni n. sp. from diamond stingray Hypanus dipterurus (Jordan & Gilbert) is differentiated from congeners by a combination of characters including total length of the whole worms, size of the scolex and bothridia, length of the bases of the hooks and abaxial prongs, and the size of the cirrus sacs in mature proglottids. Acanthobothrium hypanus n. sp. and A. sinaloaensis n. sp. are described from longtail stingray Hypanus longus (Garman). Acanthobothrium hypanus n. sp. is differentiated by a combination of characters including size of the scolex and bothridia, diameter of the accessory sucker, length of the abaxial prongs, size of the cirrus sacs and testes in mature proglottids. Acanthobothrium sinaloaensis n. sp. is differentiated by a combination of characters including size of the scolex and bothridia, diameter of the accessory sucker, size of the cirrus sacs in mature proglottids, number of immature proglottids, number of testes, the length of the lobes of the ovary and the absence of velum between medial margin of bothridia. Thirteen species of Acanthobothrium have been reported previously from elasmobranchs from the Pacific coast of Mexico. In the present study, the number of species is elevated to 16.


PLEASE send your new papers to
juergen.pollerspoeck@shark-references.com or 

Latest Research Articles

Extant Chondrichthyes:
ADAMIDOU, A. & PARDALOU, A. & TSIKLIRAS, A.C. (2020) Length-Weight Relationships of 31 Fish and Invertebrate Species in the Northern Aegean Sea (Eastern Mediterranean Sea). Thalassas, in press https://dx.doi.org/10.1007/s41208-020-00207-x
BEZERRA, N.P.A. & MACENA, B.C.L. & TRAVASSOS, P. & AFONSO, P. & HAZIN, F.H.V. (2020) Evidence of site fidelity and deep diving behaviour of scalloped hammerhead shark (Sphyrna lewini) around the Saint Peter and Saint Paul Archipelago, in the equatorial Mid-Atlantic ridge. Marine and Freshwater Research, 71 (6): 708-718 https://dx.doi.org/10.1071/MF19029
BEZERRA, N.P.A. & MACENA, B.C.L. & TRAVASSOS, P. & AFONSO, P. & HAZIN, F.H.V. (2020) Corrigendum to: Evidence of site fidelity and deep diving behaviour of scalloped hammerhead shark (Sphyrna lewini) around the Saint Peter and Saint Paul Archipelago, in the equatorial Mid-Atlantic ridge. Marine and Freshwater Research, 71 (6): 719 - 721 https://dx.doi.org/10.1071/MF19029_CO
BOWLBY, H.D. & GIBSON, A.J.F. (2020) Implications of life history uncertainty when evaluating status in the Northwest Atlantic population of white shark (Carcharodon carcharias). Ecology and Evolution, in press https://dx.doi.org/10.1002/ece3.6252
BYRNES, E.E. & LEAR, K.O. & MORGAN, D.L. & GLEISS, A.C. (2020) Respirometer in a box: development and use of a portable field respirometer for estimating oxygen consumption of large-bodied fishes. Journal of Fish Biology, 96 (4): 1045-1050 https://dx.doi.org/10.1111/jfb.14287
CACERES, A. & SCHLOBIN, N.A. & LAM, S. & ZAMORA, J. & SEGURA, J.L. (2020) Stingray spear injury to the pediatric spinal cord: case report and review of the literature. Childs Nervous System, in press https://dx.doi.org/10.1007/s00381-020-04629-z
CHATZISPYROU, A. & GUBILI, C. & LAIAKI, M. & MANTOPOULOU-PALOUKA, D. & KAVADAS, S. (2020) First record of the marbled ray, Dasyatis marmorata (Elasmobranchii: Dasyatidae), from Greece (central Aegean Sea). Biodiversity Data Journal, 8: E51100 https://dx.doi.org/10.3897/BDJ.8.e51100
COLEFAX, A.P. & KELAHER, B.P. & PAGENDAM, D.E. & BUTCHER, P.A. (2020) Assessing White Shark (Carcharodon carcharias) Behavior Along Coastal Beaches for Conservation-Focused Shark Mitigation. Frontiers in Marine Science, 7: 268 https://dx.doi.org/10.3389/fmars.2020.00268
DA SILVA, G.A. & POSCAI, A.N. & DA SILVA CASAS, A.L. (2020) Injuries Caused by Freshwater Stingrays in the Western Amazon: Folk Medicine and Beliefs. Ethnobiology Letters, 11 (1): 1-13 https://dx.doi.org/10.14237/ebl.11.1.2020.1586
DOAN, M.D. & KAJIURA, S.M. (2020) Adult blacktip sharks (Carcharhinus limbatus) use shallow water as a refuge from great hammerheads (Sphyrna mokarran). Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14342
DOLGANOV, V.N. (2020) A Morphological Description and the Biology of Poorly Known Deep-Sea Skates of the Genus Bathyraja Ishiyama, 1958 (Rajidae) from the Northwestern Pacific Ocean. Russian Journal of Marine Biology, 46 (2): 73–80 https://dx.doi.org/10.1134/S1063074020020030
EVE, O. & MATZ, H. & DOOLEY, H. (2020) Proof of long-term immunological memory in cartilaginous fishes. Developmental and Comparative Immunology, 108: 103674 https://dx.doi.org/10.1016/j.dci.2020.103674
FELDHEIM, K.A. & JABADO, R.W. & CHAPMAN, D.D. & CARDE?OSA, D. & MADDOX, J.D. (2020) Microsatellite primer development in elasmobranchs using next generation sequencing of enriched libraries. Molecular Biology Reports, 47 (4): 2669-2675 https://dx.doi.org/10.1007/s11033-020-05357-y
FIELDS, A.T. & FISCHER, G.A. & SHEA, S.K.H. & ZHANG, H. & FELDHEIM, K.A. & CHAPMAN, D.D. (2020) DNA Zip-coding: identifying the source populations supplying the international trade of a critically endangered coastal shark. Animal Conservation, in press https://dx.doi.org/10.1111/acv.12585
GOMES-DOS-SANTOS, A. & ARRONDO, N.V. & MACHADO, A.M. & VERISSIMO, A. & PEREZ, M. & ROMAN, E. & CASTRO, L.F.C. & FROUFE, E. (2020) The complete mitochondrial genome of the deep-water cartilaginous fish Hydrolagus affinis (de Brito Capello, 1868) (Holocephali: Chimaeridae). Mitochondrial DNA Part B-Resources, 5 (2): 1810-1812 https://dx.doi.org/10.1080/23802359.2020.1749154
HORNE, C.R. & HIRST, A.G. & ATKINSON, D. (2020) Selection for increased male size predicts variation in sexual size dimorphism among fish species. Proceedings of the Royal Society B-Biological Sciences, 287 (1918): 20192640 https://dx.doi.org/10.1098/rspb.2019.2640
HUANG, Q.G. & ZHANG, D. & PAN, G. (2020) Computational Model Construction and Analysis of the Hydrodynamics of a Rhinoptera javanica. Ieee Access, 8: 30410-30420 https://dx.doi.org/10.1109/access.2020.2970942
IKI, A. & ANDERSON, W.G. & DECK, C.A. & OGIHARA, M.H. & IKEBA, K. & KATAOKA, H. & HYODO, S. (2020) Measurement of 1 alpha hydroxycorticosterone in the Japanese banded houndshark, Triakis scyllium, following exposure to a series of stressors. General and Comparative Endocrinology, 292: 113440 https://dx.doi.org/10.1016/j.ygcen.2020.113440
IZZO, C. & GILLANDERS, B.M. (2020) Port Jackson Shark Growth Is Sensitive to Temperature Change. Frontiers in Marine Science, 7: 240 https://dx.doi.org/10.3389/fmars.2020.00240
JAYASANKAR, A.K. & SEIDEL, R. & HOSNY, A. & WEAVER, J.C. & FRATZL, P. & CHEN, J. & DEAN, M.N. (2020) Multi-scale modeling and mechanical performance characterization of stingray skeleton-inspired tessellations. Journal of the Mechanics and Physics of Solids, 138: 103906 https://dx.doi.org/10.1016/j.jmps.2020.103906
KOUSTENI, V. & MEGALOFONOU, P. (2020) Reproductive strategy of Scyliorhinus canicula (L., 1758): a holistic approach based on macroscopic measurements and microscopic observations of the reproductive organs. Marine and Freshwater Research, 71 (6): 596-616 https://dx.doi.org/10.1071/MF18474
KOVACIC, M. & LIPEJ, L. & DULCIC, J. (2020) Evidence approach to checklists: critical revision of the checklist of the Adriatic Sea fishes. Zootaxa, 4767 (1): 1-55 https://dx.doi.org/10.11646/zootaxa.4767.1.1
LEHMAN, R.N. & POULAKIS, G.R. & SCHARER, R.M. & SCHWEISS, K.E. & HENDON, J.M. & PHILLIPS, N.M. (2020) An environmental DNA tool for monitoring the status of the Critically Endangered Smalltooth Sawfish, Pristis pectinata, in the western Atlantic. Conservation Genetics Resources, in press https://dx.doi.org/10.1007/s12686-020-01149-5
LIU, K.M. & LIN, Y.H. & CHANG, S.K. & JOUNG, S.J. & SU, K.Y. (2020) Examining an ontogenetic shift in the diet of the whitespotted bamboo shark Chiloscyllium plagiosum in northern Taiwanese waters. Regional Studies in Marine Science, 35: 101234 https://dx.doi.org/10.1016/j.rsma.2020.101234
LÓPEZ-ROMERO, F.A. & KLIMPFINGER, C. & TANAKA, S. & KRIWET, J. (2020) Growth trajectories of prenatal embryos of the deep‐sea shark Chlamydoselachus anguineus (Chondrichthyes). Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14352
MAES, T. & DE JEL, J.V. & VETHAAK, A.D. & DESENDER, M. & BENDELL, V.A. & VAN VELZEN, M. & LESLIE, H.A. (2020) You Are What You Eat, Microplastics in Porbeagle Sharks From the North East Atlantic: Method Development and Analysis in Spiral Valve Content and Tissue. Frontiers in Marine Science, 7: 273 https://dx.doi.org/10.3389/fmars.2020.00273
MANCUSI, C. & BAINO, R. & FORTUNA, C. & DE SOLA, L. & MOREY, G. & BRADAI, M. & KALLIANOTIS, A. & SOLDO, A. & HEMIDA, F. & SAAD, A. & DIMECH, M. & PERISTERAKI, P. & BARICHE, M. & CLÒ, S. & DE SABATA, E. & CASTELLANO, L. & GARIBALDI, F. & LANTERI, L. & TINTI, F. & PAIS, A. & SPERONE, E. & MICARELLI, P. & POISSON, F. & SION, L. & CARLUCCI, R. & CEBRIAN-MENCHERO, D. & SÉRET, B. & FERRETTI, F. & EL-FAR, A. & SAYGU, I. & SHAKMAN, E. & BARTOLI, A. & GUALLART, J. & DAMALAS, D. & MEGALOFONOU, P. & VACCHI, M. & BOTTARO, M. & NOTARBARTOLO DI SCIARA, G. & FOLLESA, M. & CANNAS, R. & KABASAKAL, H. & ZAVA, B. & CAVLAN, G. & JUNG, A. & ABUDAYA, M. & KOLITARI, J. & BARASH, A. & JOKSIMOVIC, A. & MARČETA, B. & GONZALEZ VILAS, L. & TIRALONGO, F. & GIOVOS, I. & BARGNESI, F. & LELLI, S. & BARONE, M. & MORO, S. & MAZZOLDI, C. & CHARIS, C. & ABELLA, A. & SERENA, F. (2020) MEDLEM database, a data collection on large Elasmobranchs in the Mediterranean and Black seas. Mediterranean Marine Science, 0: 276-288 https://dx.doi.org/10.12681/mms.21148
MARONGIU, M.F. & PORCU, C. & BELLODI, A. & CANNAS, R. & CARBONARA, P. & CAU, A. & COLUCCIA, E. & MOCCIA, D. & MULAS, A. & PESCI, P. & FOLLESA, M.C. (2020) Abundance, distribution and reproduction of the Data-Deficient species (Squalus blainville) around Sardinia Island (central western Mediterranean Sea) as a contribution to its conservation. Marine and Freshwater Research, in press https://dx.doi.org/10.1071/MF19372
MARTINS, M.F. & ROLIM, F.A. & GADIG, O.F.B. (2020) Report on fetal mummification in the scalloped hammerhead shark Sphyrna lewini. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14368
MASON, J.G. & ALFARO-SHIGUETO, J. & MANGEL, J.C. & CROWDER, L.B. & ARDOIN, N.M. (2020) Fishers' solutions for hammerhead shark conservation in Peru. Biological Conservation, 243: Unsp 108460 https://dx.doi.org/10.1016/j.biocon.2020.108460
MEYER, L. & WHITMARSH, S.K. & NICHOLS, P.D. & REVILL, A.T. & HUVENEERS, C. (2020) The effects of wildlife tourism provisioning on non-target species. Biological Conservation, 241: Unsp 108317 https://dx.doi.org/10.1016/j.biocon.2019.108317
MOLINA, J.M. & FINOTTO, L. & WALKER, T.I. & REINA, R.D. (2020) The effect of gillnet capture on the metabolic rate of two shark species with contrasting lifestyles. Journal of Experimental Marine Biology and Ecology, 526: 151354 https://dx.doi.org/10.1016/j.jembe.2020.151354
MORICK, D. & DAVIDOVICH, N. & BIGAL, E. & ROSENBLUTH, E. & BOUZNACH, A. & ROKNEY, A. & RON, M. & WOSNICK, N. & TCHERNOV, D. & SCHEININ, A.P. (2020) Fatal Infection in a Wild Sandbar Shark (Carcharhinus plumbeus), Caused by Streptococcus agalactiae, Type Ia-ST7. Animals, 10 (2): 284 https://dx.doi.org/10.3390/ani10020284
MUSTIKA, P.L.K. & ICHSAN, M. & BOOTH, H. (2020) The Economic Value of Shark and Ray Tourism in Indonesia and Its Role in Delivering Conservation Outcomes. Frontiers in Marine Science, 7: 261 https://dx.doi.org/10.3389/fmars.2020.00261
NIELLA, Y. & SMOOTHEY, A.F. & PEDDEMORS, V. & HARCOURT, R. (2020) Predicting changes in distribution of a large coastal shark in the face of the strengthening East Australian Current. Marine Ecology Progress Series, 642: 163-177 https://dx.doi.org/10.3354/meps13322
OTWAY, N. (2020) Ultrasound-guided sampling of the lateral abdominal vein in the grey nurse shark (Carcharias taurus, Rafinesque 1810). Veterinary Medicine and Science, in press
PALACIOSHERNÁNDEZ, D. & CASTILLOGÉNIZ, J.L. & MÉNDEZLOEZA, I. & PÉREZJIMÉNEZ, J.C. (2020) Temporal and latitudinal comparisons of reproductive parameters in a heavily exploited shark, the bonnethead, Sphyrna tiburo (L. 1758), in the southern Gulf of Mexico. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14330
SABADIN, D.E. & LUCIFORA, L.O. & BARBINI, S.A. & FIGUEROA, D.E. & KITTLEIN, M. (2020) Towards regionalization of the chondrichthyan fauna of the Southwest Atlantic: a spatial framework for conservation planning. ICES Journal of Marine Science, in press: fsaa064 https://dx.doi.org/10.1093/icesjms/fsaa064
SANDOVAL-CASTILLO, J. & BEHEREGARAY, L.B. (2020) Oceanographic heterogeneity influences an ecological radiation in elasmobranchs. Journal of Biogeography, in press https://dx.doi.org/10.1111/jbi.13865
SELLHEIM, N. (2020) The CITES appendix II-Listing of mako sharks - Revisiting counter arguments. Marine Policy, 115: 103887 https://dx.doi.org/10.1016/j.marpol.2020.103887
SHERMAN, C.S. & HEUPEL, M.R. & MOORE, S.K. & CHIN, A. & SIMPFENDORFER, C.A. (2020) When sharks are away, rays will play: effects of top predator removal in coral reef ecosystems. Marine Ecology Progress Series, 641: 145-157 https://dx.doi.org/10.3354/meps13307
SILKIN, Y.A. & SILKINA, E.N. & SILKIN, M.Y. (2020) A Study of Some Biochemical Properties of Ecto-ATPase (Nucleotidase) in Erythrocytes of the Black Sea Thornback Ray Raja clavata L. Journal of Evolutionary Biochemistry and Physiology, 56 (1): 31-40 https://dx.doi.org/10.1134/s0022093020010044
SINGH, C.M.L. & AGUIAR-SANTOS, J. & FERREIRA, E.J.G. & EVARISTO, E.D. & FREITAS, C.E.D. (2020) Spatial and Temporal Distribution of a Multiple Gear Fishing Fleet Exploiting the Caribbean Sea and North Brazil Shelf Large Marine Ecosystems. Marine and Coastal Fisheries, 12 (2): 100-112 https://dx.doi.org/10.1002/mcf2.10113
SIU, S.H. & LIU, S.Y.V. & LIU, K.M. & TSAI, W.P. (2020) Development and characterization of novel microsatellite loci for an endangered hammerhead shark Sphyrna lewini by using shotgun sequencing. Taiwania, 65 (2): 261-263 https://dx.doi.org/10.6165/tai.2020.65.261
SMITH, M. & MANZANARES, E. & UNDERWOOD, C. & HEALY, C. & CLARK, B. & JOHANSON, Z. (2020) Holocephalan (Chondrichthyes) dental plates with hypermineralized dentine as a substitute for missing teeth through developmental plasticity. Journal of Fish Biology, in press https://dx.doi.org/10.1111/jfb.14302
STOECKLE, M.Y. & DAS MISHU, M. & CHARLOP-POWERS, Z. (2020) Improved Environmental DNA Reference Library Detects Overlooked Marine Fishes in New Jersey, United States. Frontiers in Marine Science, 7: 226 https://dx.doi.org/10.3389/fmars.2020.00226
TAMBURIN, E. & ELORRIAGA-VERPLANCKEN, F.R. & ESTUPINAN-MONTANO, C. & MADIGAN, D.J. & SANCHEZ-GONZALEZ, A. & PADILLA, M.H. & WCISEL, M. & GALVAN-MAGANA, F. (2020) New insights into the trophic ecology of young white sharks (Carcharodon carcharias) in waters off the Baja California Peninsula, Mexico. Marine Biology, 167 (5 https://dx.doi.org/10.1007/s00227-020-3660-8
TEIXEIRA, G. & RAIMUNDO, J. & GOULART, J. & COSTA, V. & MENEZES, G.M. & CAETANO, M. & PACHECO, M. & MARTINS, I. (2020) Hg and Se composition in demersal deep-sea fish from the North-East Atlantic. Environmental Science and Pollution Research, in press https://dx.doi.org/10.1007/s11356-020-08970-3
THOMAZI, G.O.C. & COSTA, A. & RODRIGUES, J.P. & ALVES, G.J. & NETO, J.P.P. & TURIBIO, T.D. & ROCHA, A.M. & AIRES, R.D. & SEIBERT, C.S. & SPENCER, P.J. & GALISTEO, A.J. & DE ANDRADE, H.F. & DO NASCIMENTO, N. (2020) Paratrygon aiereba irradiated anti-mucus serum reduce edematogenic activity induced in experimental model. Toxicon, 178: 13-19 https://dx.doi.org/10.1016/j.toxicon.2020.02.012
TINHAN, T.C. & O'LEARY, S.J. & PORTNOY, D.S. & ROOKER, J.R. & GELPI, C.G. & WELLS, R.J.D. (2020) Natural tags identify nursery origin of a coastal elasmobranch Carcharhinus leucas. Journal of Applied Ecology, in press https://dx.doi.org/10.1111/1365-2664.13627
TOLOTTI, M.T. & FORGET, F. & CAPELLO, M. & FILMALTER, J.D. & HUTCHINSON, M. & ITANO, D. & HOLLAND, K. & DAGORN, L. (2020) Association dynamics of tuna and purse seine bycatch species with drifting fish aggregating devices (FADs) in the tropical eastern Atlantic Ocean. Fisheries Research, 226: 105521 https://dx.doi.org/10.1016/j.fishres.2020.105521
VOGLER, R. & GONZALEZ, C. & SEGURA, A.M. (2020) Spatio-temporal dynamics of the fish community associated with artisanal fisheries activities within a key marine protected area of the Southwest Atlantic (Uruguay). Ocean & Coastal Management, 190: 105175 https://dx.doi.org/10.1016/j.ocecoaman.2020.105175
WEISBROD, T.C. & WALSH, M.T. & MARQUARDT, S. & GIGLIO, R.F. (2020) Computed Tomography Diagnosis of Pneumothorax and Cardiac Foreign Body Secondary to Stingray Injury in a Bottlenose Dolphin (Tursiops truncatus). Aquatic Mammals, 46 (3): 326-330 https://dx.doi.org/10.1578/AM.46.3.2020.326
WHEELER, C.R. & GERVAIS, C.R. & JOHNSON, M.S. & VANCE, S. & ROSA, R. & MANDELMAN, J.W. & RUMMER, J.L.  (2020) Anthropogenic stressors influence reproduction and development in elasmobranch fishes. Reviews in Fish Biology and Fisheries, 30: 373–386 https://dx.doi.org/10.1007/s11160-020-09604-0
WHITE, W.T. & BAJE, L. & APPLEYARD, S.A. & CHIN, A. & SMART, J.J. & SIMPFENDORFER, C.A. (2020) Shark longline fishery of Papua New Guinea: size and species composition and spatial variation of the catches. Marine and Freshwater Research, 71 (6): 627-640 https://dx.doi.org/10.1071/MF19191
WHITEHEAD, D.A. & MURILLO-CISNEROS, D. & ELORRIAGA-VERPLANCKEN, F.R. & HACOHEN-DOMENE, A. & DE LA PARRA, R. & GONZALEZ-ARMAS, R. & GALVAN-MAGANA, F. (2020) Stable isotope assessment of whale sharks across two ocean basins: Gulf of California and the Mexican Caribbean. Journal of Experimental Marine Biology and Ecology, 527: 151359 https://dx.doi.org/10.1016/j.jembe.2020.151359
WOO, M. & SEOL, B.G. & KANG, K.H. & CHOI, Y.H. & CHO, E.J. & NOH, J.S. (2020) Effects of collagen peptides from skate (Raja kenojei) skin on improvements of the insulin signaling pathway via attenuation of oxidative stress and inflammation. Food & Function, 11 (3): 2017-2025 https://dx.doi.org/10.1039/c9fo02667c
YOUNG, C.N. & CARLSON, J.K. (2020) The biology and conservation status of the oceanic whitetip shark (Carcharhinus longimanus) and future directions for recovery. Reviews in Fish Biology and Fisheries, 30: 293–312 https://dx.doi.org/10.1007/s11160-020-09601-3
ZHAO, C.C. & EUN, J.B. (2020) Structural and Ultrastructural Changes in Muscle Tissue of Skate (Raja kenojel) during Fermentation at 10 degrees C. Journal of Aquatic Food Product Technology, 29 (5): 486-492 https://dx.doi.org/10.1080/10498850.2020.1748780

Extinct Chondrichthyes:

DENTZIEN-DIAS, P. & HUNT, A.P. & LUCAS, S.G. & FRANCISCHINI, H. & GULOTTA, M. (2020) Coprolites from shallow marine deposits of the Nanjemoy Formation, Lower Eocene of Virginia, USA. Lethaia, in press https://dx.doi.org/10.1111/let.12380
LASSERON, M. & ALLAIN, R. & GHEERBRANT, E. & HADDOUMI, H. & JALIL, N.E. & METAIS, G. & RAGE, J.C. & VULLO, R. & ZOUHRI, S. (2020) New data on the microvertebrate fauna from the Upper Jurassic or lowest Cretaceous of Ksar Metlili (Anoual Syncline, eastern Morocco). Geological Magazine, 157 (3): 367-392 https://dx.doi.org/10.1017/s0016756819000761
NERAUDEAU, D. & VULLO, R. & BENEFICE, P. & BRETON, G. & DEPRE, E. & GASPARD, D. & GIRARD, V. & LE COULS, M. & MOREAU, J.D. & NEL, A. & PERRICHOT, V. & SOLORZANO-KRAEMER, M.M. & WAPPLER, T. (2020) The paralic Albian-Cenomanian Puy-Puy Lagerstatte (Aquitaine Basin, France): An overview and new data. Cretaceous Research, 111: 104124 https://dx.doi.org/10.1016/j.cretres.2019.03.022
SLATER, T.S. & ASHBROOK, K. & KRIWET, J. (2020) Evolutionary relationships among bullhead sharks (Chondrichthyes, Heterodontiformes). Papers in Palaeontology, in press https://dx.doi.org/10.1002/spp2.1299
TRIKOLIDI, F.A. & NOVIKOV, I.V. (2020) The First Find of the Dental Plate of the Genus Myliobatis Cuvier (Elasmobranchii: Batomorphii) in the Upper Paleocene of the Crimea. Paleontological Journal, 54 (2): 166-170 https://dx.doi.org/10.1134/s0031030120020148
VILLAFAÑA, J.A. & HERNANDEZ, S. & ALVARADO, A. & SHIMADA, K. & PIMIENTO, C. & RIVADENEIRA, M.M. & KRIWET, J. (2020) First evidence of a palaeo-nursery area of the great white shark. Scientific Reports, 10: 8502 https://dx.doi.org/10.1038/s41598-020-65101-1
VILLAFAÑA, J.A. & MARRAMÀ, G. & KLUG, S. & POLLERSPÖCK, J. & BALSBERGER, M. & RIVADENEIRA, M. & KRIWET, J. (2020) Sharks, rays and skates (Chondrichthyes, Elasmobranchii) from the Upper Marine Molasse (middle Burdigalian, early Miocene) of the Simssee area (Bavaria, Germany), with comments on palaeogeographic and ecological patterns. Paläontologische Zeitschrift, in press  https://dx.doi.org/10.1007/s12542-020-00518-7


BRAY, R.A. & WAESCHENBACH, A. & LITTLEWOOD, D.T.J. & HALVORSEN, O. & OLSON, P.D. (2020) Molecular circumscription of new species of Gyrocotyle Diesing, 1850 (Cestoda) from deep-sea chimaeriform holocephalans in the North Atlantic. Systematic Parasitology, 97 (3): 285-296 https://dx.doi.org/10.1007/s11230-020-09912-w
DENNIS, M.M. & YEOW, J.W.A. & NICOLACI, B. & SAMPLE, S.H. & FREEMAN, M.A. (2020) Meningitis associated with dracunculid nematodes in a spotted eagle ray (Aetobatus narinari). Journal of Fish Diseases, in press https://dx.doi.org/10.1111/jfd.13160
SHAMSI, S. & BARTON, D.P. & ZHU, X.C. (2020) Description and genetic characterisation of Pulchrascaris australis n. sp. in the scalloped hammerhead shark, Sphyrna lewini (Griffin & Smith) in Australian waters. Parasitology Research, in press https://dx.doi.org/10.1007/s00436-020-06672-w
ZARAGOZA-TAPIA, F. & PULIDO-FLORES, G. & MONKS, S. (2020) Three new species of Acanthobothrium Blanchard, 1848 (Cestoda: Onchoproteocephalidea) in Stingrays (Dasyatidae) from the Pacific coast in Mexico. Zootaxa, 4766 (1): 157-172  https://dx.doi.org/10.11646/zootaxa.4766.1.8


Little skates could hold the key to cartilage therapy in humans

Date: May 12, 2020
Source: Marine Biological Laboratory
Summary: Unlike humans and other mammals, the skeletons of sharks, skates, and rays are made entirely of cartilage and they continue to grow that cartilage throughout adulthood. New research finds that adult skates go one step further than cartilage growth: They can also spontaneously repair injured cartilage. This is the first known example of adult cartilage repair in a research organism. The team also found that newly healed skate cartilage did not form scar tissue.

Can't touch this! Video shows blacktip sharks use shallow water to flee huge predators

Aerial video provides first evidence of adult sharks using shallow water to escape the great hammerhead

Date: May 13, 2020
Source: Florida Atlantic University
Summary: Aerial drone footage provides the first evidence of adult blacktip sharks using shallow waters as a refuge from a huge predator -- the great hammerhead. Before this study, documentation of adult sharks swimming in shallower waters to avoid predation did not exist. Unmanned aerial vehicles enable scientists to unobtrusively observe behaviors in the wild, providing insight into seldom-seen predator-prey interactions. When it comes to sharks, this 'hammerhead' time video proves you 'can't touch this.'


First fossil nursery of the great white shark discovered

Paleo-kindergarten ensured evolutionary success millions of years ago

Date: May 22, 2020
Source: University of Vienna
Summary: An international research team discovered the first fossil nursery area of the great white shark, Carcharodon carcharias in Chile. This discovery provides a better understanding of the evolutionary success of the largest top predator in today's oceans in the past and could contribute to the protection of these endangered animals.