NEWSLETTER 09/2019 09.09.2019

Please acknowledge use of the database www.shark-references.com in your publications, and cite: 

Pollerspöck, J. & Straube, N. 2019, Bibliography database of living/fossil sharks, rays and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali), www.shark-references.com, World Wide Web electronic publication, Version 2019

New open access paper from Team Shark-references:

In addition to articulated, mostly formaldehyde-fixed and ethanol-preserved, taxidermy or anatomical specimens, sharks and rays are represented in scientific collections by numerous jaws and isolated teeth. These specimens often source from historical collections where existing informations about species, sex or geographic origin in many cases are scarce, incomplete or incorrect. The identification key for jaws and teeth presented herein focuses on squalomorph sharks, which comprise almost 34 % of all sharks with 179 species in 31 genera and 11 families. The key is essentially based on the following characters: vascularisation stage, labial apron, number of cusplets, distal heel, lingual ornamentation, cutting edge, and dentition kind. The key allows the identification to genus level. It is further supplemented by a comprehensive glossary of tooth morphological terms as well as an updated checklist of all currently described squalomorph sharks with indication of the distribution and the dental formula.

POLLERSPÖCK, J. & STRAUBE, N. 2019 An identification key to elasmobranch genera based on dental morphological characters Part A: Squalomorph sharks (Superorder Squalomorphii). Bulletin of Fish Biology, 18 (1/2): 77-105

free download via research gate



Would you like to become a shark-reference partner? Please contanct us per E-mail!



Partner in Google-Maps




New Images

Many thanks to the following people for providing images:

Frederik Mollen (Elasmobranch Research Belgium) for the images of Atlantoraja castelnaui (MIRANDA RIBEIRO,1907), ERB 0553, female, 95,5 DW, 135,5 TL, Uruguay:

Many thanks to Jean-Francois LHOMME for the images of Heterodontus vincenti (LERICHE, 1905), Eocene, Lutetian, Paris  Basin, France:

Balanov A.A., The head Laboratory of Ichthyology, IBM FEB RAS, Vladivostok, Russia for the image of the holotye of Etmopterus parini DOLGANOV & BALANOV, 2018:

Kelsi M. Rutledge for images of the new described Pseudobatos buthi RUTLEDGE, 2019:

The Team of iSea Greece (https://www.facebook.com/iSea.org/) for images of Hexanchus griseus (BONNATERRE, 1788):

Mark Sabaj Perez for images of the paratype of Mustelus minicanis HEEMSTRA, 1997:


Missing papers:

Many thanks to all friends of shark-references, who sent us some missing papers last month!

Shark-References would kindly like to ask you for your contribution to this project.

At the moment we are looking for some of the following papers:

Extinct Chondrichthyes:

MATSUMOTO, H. (1936) Upper Miocene vertebrates from Kumanodô, Natori district, province of Rikuzen. Dobutsugaku Zasshi, 48: 475–480, 5 fig.

DERANIYAGALA, P.E.P. (1937) Some Miocene fishes from Ceylon. Ceylon Journal of Science, 20 (3): 355–367,9 fig.

KUDRIN, L.N. (1957) O nachodkach zubov akul v otlozenijach miocena jugozapadnoj okrainy Russkoj platformy (On some finds of the shark's teeth in the Miocene's deposits in the South-Western border of the Russian platform). Geol. Sborn. Lvov. Geol. Obsc. (Geol. J . of Geol. Soc. in Lvov), 4, Lvov.

BELTAN, L. (1959) Sur la présence du genre Carcharodon Müller et Henle dans le Miocène de la Martinique (Petites Antilles francaises). C.R. Sommaire des Seances de la Societe Geologique de France, 6: 156–158

ZBYSZEWSKI, G. & FERREIRA, O. DA V. (1962) La faune Miocène de l’île de Santa Maria (Açores). Comunicações dos Serviços Geológicos de Portugal, 46: 247–289

MENESINI, E. (1967) I Pesci miocenici delle Arenarie di Ponsano. Atti della Società Toscana di Scienze Naturali, Serie A, 74 (1): 1–22.

DERANIYAGALA, P.E.P. (1969) A Miocene vertebrate faunule from the Malu Member of Ceylon. Spolia Zeylan, 31: 551–570, 19 fig., 10 tabl.

CAPPETTA, H. (1969) Les gisements de vertébrés de la région montpelliéraine. 2. Gisements miocènes. Bulletin du Bureau de Recherches géologiques et minières, 2 (1): 19–30

COMASCHI CARIA, I. (1973) I pesci del Miocene della Sardigna. Stabilimento Tipografico Editoriale Fossataro, Cagliari. 39 pp.

ALVINERIE, J. & ANDREIEFF, P. & ANGLADA, R. & AUBERT, J. & CAPPETTA, H. & CARALP, M. & CARATINI, C. & CARBONNEL, G. & CATZIGRAS, F. & COURME-RAULT, M.-D. & CHATEAUNEUF, J.-J. & DEMARCQ, G. & DUCASSE, O. & FATTON, E. & GLAÇON, G. & LABRACHERIE, M. & LAURIAT, A. & LE CALVEZ, Y. & LORENZ, C. & MAGNE, J. & MARGEREL, J.-P. & POIGNANT, A. & PUJOL, C. & ROGER, J. & ROMAN, J. & BLONDEAU, A. & MULLER, C. (1973) A propos de la limite oligo-miocène: résultats préliminaires d'une recherche collective sur les gisements d'Escornébéou (Saint-Géours-de-Maremne, Landes, Aquitaine méridionale). Présence de Globigerinoides dans les faunes de l'Oligocène supérieur. Comptes rendus sommaires des séances de la Société géologique de France: 75–76

BALBINO, A.C. (1996) Sharks from the Middle and early Upper Miocene from Lisbon, Portugal. A check-list. Comunicações do Instituto Geológico e Mineiro, 82: 141–144

KOZLOV, V.A. (2001) (A new species of carcharhinid shark from the Lower Miocene sediments of Northern Aral region) «in russian». Materialy po Stratigrafii i Paleontologii Urala, 6: 92–95

BOYD, B.M. (2016) Fossil sharks and rays of Gainesville creeks; Alachua County, Florida: Hogtown group; (middle Miocene to lower Pliocene). Florida Paleontological Society, Special Publication

Extant Chondrichthyes:

DEVINCENZI, G.J. & TEAGUE, G.W. (1942) Ictiofauna del Rio Uruguay medio. Anales del Museo Nacional de Historia Natural de Montevideo, (Serie 2), 5 ( 4): 1–100 + index + i–viii, Pls. 1–6

KAMOHARA, T. (1943) Some unrecorded and two new fishes from Prov. Tosa, Japan. Bulletin of the Biogeographical Society of Japan, 13 (17): 125–137

DE BUEN, F. (1950) Contribuciones a la Ictiología. II. El tiburón vitamínico de la costa uruguaya Galeorhinus vitaminicus nov. sp., y algunas consideraciones generales sobre su biología. Publicaciones Cientificas, Servicio Oceanografico y de Pesca, Ministerio de Industrias y Trabajo, Montevideo No. 4: 153–162.

CADENAT, J. (1951) Initiations Africaines. III. Poissons de Mer du Sénégal. Institute Francais d'Afrique Noire. Initiations Africaines. III. Poissons de Mer du Sénégal.: 1–345

DE BUEN, F. (1952) El tiburón vitamínico de la costa Uruguaya, Galeorhinus vitaminicus nov. sp. y algunas consideraciones generales sobre su biologia. Rev. Fac.Hum.Cienc.Univ.Repúbl.Montevid., 7: 87–116

WEIBEZAHN, F.H. (1953) Una nueva especie de Scyliorhinus de Venezuela (Chondrichthyes - Elasmobranchii). Novedades cientificas. Serie zoológica. Museo de Historia Natural La Salle, 9: 1–7.

SMITH, J.L.B. (1958) The mystery killer, the new shark Carcharhinus vanrooyeni. Veld & Vlei, 3 (9): 12–14, 28.

SICCARDI, E. (1961) Cetorhinus en el Atlantico sur (Elasmobranchii: Cetorhinidae). Actas y trabajos del Primer Congreso Sudamericano de Zoologia, 4 (5): 251–263

CADENAT, J. (1963) Notes d'ichtyologie ouest-africaine. XXXIX. Notes sur les requins de la famille des Carchariidae et formes apparentées de l'Atlantique ouest-africain (avec la description d'une espèce nouvelle: Pseudocarcharias pelagicus, classée dans un sous-genre nouveau). Bulletin de l'Institut Français d'Afrique Noire (A), 25 (2): 526–537, 9 fig.

CASTEX, M.N. (1963) Una nueva especie de raya fluvial: Potamotrygon pauckei. Notas distintivas Boletín de la Academia Nacional de Ciencias (Córdoba), 43: 289–294

CADENAT, J. & MAUL, G.E. (1966) Note d'ichthyologie ouest-africaine. XLIII. Description d'une espèce nouvelle du genre Apristurus, A. maderensis. Bulletin de l'Institut Français d'Afrique Noire (A), 28 (2): 769–782

ABELLA, A. (1972) Halazgo de una nueva especie de Carcharinus en las costas de Rocha, Uruguay. Boletín de la Sociedad Zoológica del Uruguay, 2: 102–106

GUITART-MANDAY, D.J. (1972) Un nuevo género y especies de tiburón de la Familia Triakidae. Poeyana (Ser.A), 1972 (99): 1–4

GUBANOV, E.P. & SCHLEIB, N.A. (1980) Sharks of the Arabian Gulf. Kuwait Ministry of Public Works, Agracultural Department, Fisheries Division. Sharks of the Arabian Gulf.: 1–69

DOLGANOV, V.N. (1983) Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod. [Manual for identification of cartilaginous fishes of Far East seas of USSR and adjacent waters.] TINRO, Vladivostok. Rukovodstvo po opredeleniyu khryashchevykh ryb dal'nevostochnykh morei SSSR i sopredel'nykh vod.: 92 pp.

SHEN, S.-C. (1986) A new species of stingray Hexatrygon taiwanensis from Taiwan Strait. Journal of Taiwan Museum, 39 (1): 175–180

Please support www.shark-references.com and send missing papers (not listed papers or papers without the info-symbol) to juergen.pollerspoeck@shark-references.com or nicolas.straube@shark-references.com



Upcoming Meetings:

Annual Meeting of the Paleontological Society (Paläontologische Gesellschaft) 2019

logo_palges_201915 –18 September 2019 in Munich

We are happy to announce the Annual conference of the Paläontologische Gesellschaft (PalGes) in Munich. We aim to organize an international meeting at which cutting-edge research in the fields of palaeontology, geobiology and palaeobiology is presented. PalGes is one of the oldest and largest palaeontogical societies of the world and it is the fifth time that this conference is held in Munich (1950, 1960, 1985, 2010, 2019).

Munich is Bavaria’s beautiful capital situated at the foot of the Alps and surrounded by several picturesque lakes. It has been ranked among the top cities in terms of life quality, is famous for its museums and beer gardens. Munich is also a top place for natural sciences.

Symposia and workshops


  • Evolving ecosystems (Organizers: Alexander Nützel & Joachim Haug) more details
  • Fossil fishes in the context of evolution, environments and biogeography (Organizers: Bettina Reichenbacher, Tomas Přikryl & Gloria Arratia)
    more more details
  • Late Paleozoic and Mesozoic Plants and Floras (Organizers: Michael Krings, Benjamin Bomfleur & Christian Pott)
  • Mikropaläontologie (Organizer: Anna Pint)
  • Molecular Geobiology and Paleobiology (Organisation: Gert Wörheide, Dirk Erpenbeck, Sebastian Höhna, William Orsi) more details
  • Physiology in Deep Time (Organizers: Uwe Balthasar, Kenneth De Baets, Carl Reddin, Nussaïbah Raja Schoob) 
    more details


  • Total-evidence divergence time estimation from extinct and extant taxa (Convener: Sebastian Höhna) more details
  • Open data analysis and publication: from morphology to evolutionary patterns (Conveners: Emilia Jarochowska and Kenneth De Baets; Guest speakers: Melanie Hopkins & Stephan Lautenschlager) more details
for more information please visit


EEA Meeting 2019 in Rende


The Dipartimento di Biologia, Ecologia e Scienze della Terra – University of Calabria (DIBEST), Gruppo Italiano Ricercatori Squali, Razze e Chimere (G.R.I.S.) and the Centro Studi Squali (C.S.S.) are excited to organize the 23rd European Elasmobranch Association Meeting, which will take place in Rende (Italy) from the 16th to the 18th of October 2019.

We aim at creating a great forum where scientists, conservationists, students and all those interested in elasmobranchs will have an opportunity to share the results of their most exciting research, to look for synergies and find new collaborations.

During the conference, four plenary lectures, scientific sessions (orals and posters) and various workshops will take place. The sessions will deal with diverse topics, from evolutionary biology to ethology, from morphology to physiology, ecology and conservation. The conference will also include the Annual General Meeting of the EEA, multiple social activities, and the opportunity of exciting field underwater excursions. English will be the official conference language.

Please come back in the coming weeks to find out all the details about registration and related activities on the meeting’s website at www.dibest.unical.it/eea2019 and on the Facebook page at https://www.facebook.com/eea2019meeting/.

Provisional key dates are:

Call for abstracts and opening of registrations: 15th February 2019

End of reduced registration fees: 15th May 2019

Deadline of abstract submission: 15th June 2019

Communication of acceptance: Before August 30th



Extant Chondrichthyes:

DOLGANOV, V.N. & BALANOV, A.A. (2018):
 Etmopterus parini sp. n. (Squaliformes: Etmopteridae), a new shark species from the northwestern Pacific Ocean. Biologiya Morya (Vladivostok), 44 (6): 427-430
New species: Etmopterus parini
Abstract: A description of Etmopterus parini sp. n. (TL 376 and 341 mm), a new species of lantarn sharks caught from the epipelagic zone of the northwestern Pacific Ocean in the spring of 2010, is presented. The species is distinguished from the congeners by the following characters: body shape, body color, morphology of teeth and dermal denticles, number of trunk and caudal vertebrae, position of fins, and size and shape of luminescent markings on the flanks.

RUTLEDGE, K.M. (2019): A New Guitarfish of the Genus Pseudobatos (Batoidea: Rhinobatidae) with Key to the Guitarfishes of the Gulf of California. Copeia, 107 (3): 451-463
New species: Pseudobatos buthi
Abstract: A new guitarfish of the genus Pseudobatos is described based on 82 specimens obtained from the Gulf of California. Sixty-three morphometric measurements were taken on all specimens, and on ten specimens from each of three congeners. A principal component analysis and linear discriminant analysis were performed on these morphometric data for discrimination. The new species (∼685 mm TL) is most similar to Pseudobatos productus but differs in having a narrower maximum disc width (30–35% vs. 36–38% TL), shorter distance from nostril to disc margin (2.8–4.0% vs. 4.2–5.2% TL), narrower disc width at anterior orbit (12–19% vs. 20–22% TL), and a narrower tip of snout width (3% vs. 4–6% TL). The species is also less densely scaled between the orbits and has less pronounced rostral thorns than Pseudobatos productus. A key to the guitarfishes of the Gulf of California is also provided.

Extinct Chondrichthyes:

VILLAFAÑA, J.A. & NIELSEN, S.N. & KLUG, S. & KRIWET, J. (2019): Early Miocene cartilaginous fishes (Chondrichthyes: Holocephali, Elasmobranchii) from Chile: Diversity and palaeobiogeographic implications. Journal of South American Earth Sciences, in press
New species: Pristiophorus humboldti
Abstract: The early Miocene is characterized by warm conditions until the middle Miocene when temperatures dropped significantly. The presence of tropical to subtropical invertebrate faunas in early Miocene sediments of Chile supports the hypothesis of warm temperatures. The Neogene fossil record of chondrichthyans (holocephalans, sharks, rays and skates) has been well established for Chile. However, most studies focused on middle Miocene to Pliocene records, whereas early Miocene chondrichthyans have been rather poorly investigated up to now. The aim of this study is to describe early Miocene chondrichthyans from Chile and to discuss their paleobiogeographic and ecological implications. Here, we report seventeen chondrichthyan taxa from the early Miocene of Chile. The fauna includes the first fossil record of Mustelus from the Pacific coast of South America, the first oral tooth of Pristiophorus humboldti nov. sp. from the Neogene of South America and the first fossil record of Alopias from the Neogene of Chile. We are able to increase the total number of taxa from the early Miocene of Chile from 13 to 21. Faunal shifts in the marine waters of Chile between the early Miocene and the present reveal different biogeographical dynamics: three taxa decreased their southern latitudinal range, seven increased their southern latitudinal range, six went globally extinct and one went regionally extinct. The extinction and latitudinal changes observed in chondrichthyans can be best explained by climatic fluctuations during the Neogene and Holocene along the Pacific coast of South America. However, studies to evaluate the effect of ecological traits should be considered in the future.

ENGELBRECHT, A. & MORS, T. & REGUERO, M.A. & KRIWET, J. (2019): Skates and rays (Elasmobranchii, Batomorphii) from the Eocene La Meseta and Submeseta formations, Seymour Island, Antarctica. Historical Biology, 31 (8): 1028-1044
New genera: Marambioraja, Mesetaraja
New species: Raja manitaria, Raja amphitrita, Marambioraja leiostemma, Mesetaraja maleficapelli,
Abstract: Eocene deposits of the famous La Meseta Formation of Seymour Island, Antarctic Peninsula, yielded the most diverse Paleogene fossil elasmobranch association of the Southern Hemisphere. In this assemblage, sharks clearly dominate the fauna, whereas batoids are very rare components. Herein, we describe two new taxa of cold water tolerant skates, Marambioraja leiostemma gen. et sp. nov., and Mesetaraja maleficapelli gen. et sp. nov., two new species of the genus RajaRaja amphitrita sp. nov. and Raja manitaria sp. nov., as well as remains of warm water adapted myliobatiforms. It is, however, not possible to unambiguously assign these remains either to Myliobatidae or Rhinopteridae, or to any specific genus. Previously reported remains of Raja/Bathyrajasp. are assigned to the new described species Raja manitariasp. nov. The biogeographic distribution of extant and extinct rays and skates clearly shows that both groups are more widely distributed today than in the past, and additionally seem to have been more diverse in the Northern than the Southern Hemisphere. The occurrence, albeit rare of isolated teeth of skates (Rajidae) and rays (Myliobatidae) in the La Meseta Formation representes a minimum age constraint for their first appearance in the Southern Ocean.


SMALES, L.R. & BARTON, D.P. & CHISHOLM, L.A. (2019): Acanthocephalans from Australian elasmobranchs (Chondrichthyes) with a description of a new species in the genus Gorgorhynchus Chandler, 1934 (Rhadinorhynchidae).Systematic Parasitology, 96 (7): 565-573
New species: Gorgorhynchus occultus
AbstractGorgorhynchus occultus n. sp. is described from Sutorectus tentaculatus (Peters) (Orectolobidae) collected off Bunbury, Western Australia in 1986. The new species differs from all other species of Gorgorhynchus Chandler, 1934 by having a suite of characters including a proboscis hook formula of 18–20 rows of 8–9 hooks, a well-developed neck, irregular circles of small spines in a single anterior field, the male reproductive system limited to the posterior quarter of the trunk and three cement glands. In a survey of 284 sharks collected between 2015 and 2018 from 10 localities in Australian waters, 11 individuals were infected with acanthocephalan cystacanths. One individual of Sphyrna mokarran (Rupell) (Sphyrnidae) was infected with Corynosoma cetaceum Johnston & Best, 1931. Serrasentis sagittifer (Linton, 1889) (Rhadinorhynchidae) was found in five individuals of S. mokarran, four individuals of Syphyrna lewini (Griffith & Smith) and one individual of Carcharhinus coatesi (Whitley) (Carcharhinidae). These infections may be accidental because it has been suggested that acanthocephalans cannot tolerate the high levels of urea used by marine and esturine elasmobranchs for osmoregulation. The two most common host species examined, S. mokarran and S. lewini had the highest intensities and prevalences of infection with S. sagittifer. Although more individuals of S. lewini were examined, S. mokarranhad the higher prevalence of infection.


PLEASE send your new papers to juergen.pollerspoeck@shark-references.com or nicolas.straube@shark-references.com   

Latest Research Articles

Extant Chondrichthyes:
BARBINI, S.A. & LUCIFORA,  L.O. & SABADIN, D.E. & FIGUEROA, D.E. (2019) Ecological specialization is associated with high conservation concern in skates (Chondrichthyes, Rajiformes). Animal Conservation, in press http://dx.doi.org/10.1111/acv.12531
BORODAVKINA, M.V. & CHERNOVA, N.V. & CHEKMENEVA, N.A.  (2019) About New Findings of the Greenland Shark Somniosus microcephalus in the Kara Sea. Journal of Ichthyology, 59 (4): 623–627 http://dx.doi.org/10.1134/S0032945219030020
BROADHURST, M.K. & DOMIT, C. & TREVIZANI, T.H. & RAOULT, V. & MILLAR, R.B. (2019) Mother-embryo isotope fractionation in the pygmy devilray Mobula kuhlii cf. eregoodootenkee. Journal of Fish Biology, 95 (2): 589-593 http://dx.doi.org/10.1111/jfb.14010
CALTABELLOTTA, F.P. & SIDERS, Z.A. & MURIE, D.J. & MOTTA, F.S. & CAILLIET, G.M. & GADIG, O.B.F. (2019) Age and growth of three endemic threatened guitarfishes Pseudobatos horkelii, P. percellens and Zapteryx brevirostris in the western South Atlantic Ocean. Journal of Fish Biology, in press http://dx.doi.org/10.1111/jfb.14123
ĆETKOVIĆ, I. & PEŠIĆ, A. &  JOKSIMOVIĆ, A. & TOMANIĆ, J. & RALEVIĆ, S. (2019) Morphometric measurements of newborn blue shark Prionace glauca (Linnaeus, 1758) and characteristics of its potential parturition areas in coastal waters of Montenegro (Southeastern Adriatic). Acta Adriatica, 60 (1): 61-67 http://dx.doi.org/10.32582/aa.60.1.6
COIRATON, C. & TOVAR-AVILA, J. & GARCES-GARCIA, K.C. & RODRIGUEZ-MADRIGAL, J.A. & GALLEGOS-CAMACHO, R. & CHAVEZ-ARRENQUIN, D.A. & AMEZCUA, F. (2019) Periodicity of the growth-band formation in vertebrae of juvenile scalloped hammerhead shark Sphyrna lewini from the Mexican Pacific Ocean. Journal of Fish Biology, in press http://dx.doi.org/10.1111/jfb.14100
CRAMPTON, W.G.R. (2019) Electroreception, electrogenesis and electric signal evolution. Journal of Fish Biology, 95 (1): 92-134 http://dx.doi.org/10.1111/jfb.13922
DOCAMPO-SEARA, A. & LANOIZELET, M. & LAGADEC, R. & MAZAN, S. & CANDAL, E. & RODRIGUEZ, M.A. (2019) Mitral cell development in the olfactory bulb of sharks: evidences of a conserved pattern of glutamatergic neurogenesis. Brain Structure & Function, 224 (7): 2325-2341 http://dx.doi.org/10.1007/s00429-019-01906-9
DOLGANOV, V.N. & BALANOV, A.A. (2018) Etmopterus parini sp. n. (Squaliformes: Etmopteridae), a new shark species from the northwestern Pacific Ocean.  Biologiya Morya (Vladivostok), 44 (6): 427-430 
EL LAKHRACH, H. & HATTOUR, A. & JARBOUI, O. & BRADAI, M.N. & ESPLA, A.A.R. (2019) Spatial and temporal variations of inshore demersal fishes in the Gulf of Gabes (Tunisia, Central Mediterranean Sea). Journal of Coastal Conservation, 23 (3): 521-530 http://dx.doi.org/10.1007/s11852-019-00681-3
FAZIO, F. & IARIA, C. & SAOCA, C. & SURACI, V. & PICCIONE, G. & MARINO, F. (2019) Haematological characterization in the blackmouth catshark Galeus melastomus Rafinesque, 1810 caught in the lower Tyrrhenian Sea. Cahiers De Biologie Marine, 60 (4): 353-359  http://dx.doi.org/10.21411/cbm.a.7b66f468
FRANCIS, M.P. & MAOLAGAIN, C.O. (2019) Growth-band counts from elephantfish Callorhinchus milii fin spines do not correspond with independently estimated ages. Journal of Fish Biology, 95 (3): 743-752 http://dx.doi.org/10.1111/jfb.14060
FREITAS, R.H.A. & AGUIAR, A.A. & FREITAS, AKCHA, LIMA, S.M.Q. & VALENTIN, J.L. (2019) Unravelling the foraging behavior of the southern stingray, Hypanus americanus (Myliobatiformes: Dasyatidae) in a Southwestern Atlantic MPA. Neotropical Ichthyology, 17 (2): 180131 http://dx.doi.org/10.1590/1982-0224-20180131
GEORGE, L.W. & MARTINS, A.P.B. & HEUPEL, M.R. & SIMPFENDORFER, C.A. (2019) Fine-scale movements of juvenile blacktip reef sharks Carcharhinus melanopterus in a shallow nearshore nursery. Marine Ecology Progress Series, 623: 85-97 http://dx.doi.org/10.3354/meps13010
GERRY, S.P. & BRODEUR, L.K. & DECAPRIO, M. & KHURSIGARA, A.J. & MAZZEO, S. & NEUBAUER, D.L. (2019) Jaw muscle activation patterns of several Batoids. Environmental Biology of Fishes, 102 (9): 1193-1200 http://dx.doi.org/10.1007/s10641-019-00901-7
GLAUS, K.B.J. & BRUNNSCHWEILER, J.M. & PIOVANO, S. & MESCAM, G. & GENTER, F. & FLUEKIGER, P. & RICO, C. (2019) Essential waters: Young bull sharks in Fiji's largest riverine system. Ecology and Evolution, 9 (13): 7574-7585 http://dx.doi.org/10.1002/ece3.5304
GRACE, M.A. & HUBER, D. & TRAVIS, K. & DOOSEY, M. & FORD, J. & DECKER, S. & DENTON, J. (2019) Simulating Cookiecutter Shark bites with a 3D-printed maxillary/mandibular model. Abstract. 45th Mississippi AFS Chapter Meeting. Jackson, Mississippi 2019  
GRANT, M.I. & KYNE, P.M. & SIMPFENDORFER, C.A. & WHITE, W.T. & CHIN, A. (2019) Categorising use patterns of non-marine environments by elasmobranchs and a review of their extinction risk. Reviews in Fish Biology and Fisheries, 29 (3): 689-710 http://dx.doi.org/10.1007/s11160-019-09576-w
HUME, J.B. (2019) Higher temperatures increase developmental rate & reduce body size at hatching in the small-eyed skate Raja microocellata: implications for exploitation of an elasmobranch in warming seas. Journal of Fish Biology, 95 (2): 655-658 http://dx.doi.org/10.1111/jfb.13997
JAKOBSDOTTIR, K.B. & JONASSON, J.P. & BURGOS, J.M. & PALSSON, J. (2019) Observations of two deep-sea chimaeroids in Icelandic waters. Journal of Fish Biology, in press http://dx.doi.org/10.1111/jfb.14104
KIILU, B.K. & KAUNDA-ARARA, B. & ODDENYO, R.M. & THOYA, P. & NJIRU, J.M. (2019) Spatial distribution, seasonal abundance and exploitation status of shark species in Kenyan coastal waters. African Journal of Marine Science, 41 (2): 191-201 http://dx.doi.org/10.2989/1814232x.2019.1624614
LYONS, K. & KACEV, D. & PRETI, A. & GILLETT, D. & DEWAR, H. (2019) Organic contaminants as an ecological tool to explore niche partitioning: a case study using three pelagic shark species. Scientific Reports, 9: 12080 http://dx.doi.org/10.1038/s41598-019-48521-6
MARANDEL, F. & LORANCE, P. & TRENKEL, V.M. (2019) Determining long-term changes in a skate assemblage with aggregated landings and limited species data. Fisheries Management and Ecology, 26 (4): 365-373 http://dx.doi.org/10.1111/fme.12367
MAXWELL, S.M. & SCALES, K.L. & BOGRAD, S.J. & BRISCOE, D.K. & DEWAR, H. & HAZEN, E.L. & LEWISON, R.L. & WELCH, H. & CROWDER, L.B. (2019) Seasonal spatial segregation in blue sharks (Prionace glauca) by sex and size class in the Northeast Pacific Ocean. Diversity and Distributions, 25 (8): 1304-1317 http://dx.doi.org/10.1111/ddi.12941
MCINTURF, A.G. & STEEL, A.E. & BUCKHORN, M. & SANDSTROM, P. & SLAGER, C.J. & FANGUE, N.A. & KLIMLEY, A.P. & CAILLAUD, D. (2019) Use of a hydrodynamic model to examine behavioral response of broadnose sevengill sharks (Notorynchus cepedianus) to estuarine tidal flow. Environmental Biology of Fishes, 102 (9): 1149-1159 http://dx.doi.org/10.1007/s10641-019-00894-3
MÉNDEZ-MACÍAS, J.S. & VELÁZQUEZ-CHIQUITO, V.M. & ESTUPIÑÁN-MONTAÑO, C. & GALVÁN-MAGAÑA, F. (2019) Trophic ecology and ontogenetic shift in the diet of the sicklefin smoothhound (Mustelus lunulatus) in the southeastern Pacific Ocean. Fishery Bulletin, 117 (3): 245–257 http://dx.doi.org/10.7755/FB.117.3.11
MOORE, A.B.M. & GRUBBS, R.D. (2019) Shark and ray conservation research: Absent where the need is greatest. Aquatic Conservation-Marine and Freshwater Ecosystems, in press http://dx.doi.org/10.1002/aqc.3192
MORFIN, M. & SIMON, J. & MORANDEAU, F. & BAULIER, L.& MÉHAULT, S. & KOPP, D. (2019) Using acoustic telemetry to estimate post-release survival of undulate ray Raja undulata (Rajidae) in northeast Altantic. Ocean & Coastal Management, 178: 104848 http://dx.doi.org/10.1016/j.ocecoaman.2019.104848
MOYER, J.K. & SHANNON, S.F. & IRSCHICK, D.J. (2019) Bite performance and feeding behaviour of the sand tiger shark Carcharias taurus. Journal of Fish Biology, 95 (3): 881-892 http://dx.doi.org/10.1111/jfb.14086
MUSYL, M.K. & GILMAN, E.L. (2019) Meta-analysis of post-release fishing mortality in apex predatory pelagic sharks and white marlin. Fish and Fisheries, 20 (3): 466-500 http://dx.doi.org/10.1111/faf.12358
NEWTON, K.C. & GILL, A.B. & KAJIURA, S.M. (2019) Electroreception in marine fishes: chondrichthyans. Journal of Fish Biology, 95 (1): 135-154 http://dx.doi.org/10.1111/jfb.14068
NOSAL, A.P. & CARTAMIL, D.P. & WEGNER, N.C. & LAM, C.H. & HASTINGS, P.A. (2019) Movement ecology of young-of-the-year blue sharks Prionace glauca and shortfin makos Isurus oxyrinchus within a putative binational nursery area. Marine Ecology Progress Series, 623: 99-115 http://dx.doi.org/10.3354/meps13021
PATRICIA, F.W. & GUZMAN, D. & INIGO, B. & URTZI, I. & MARIA, B.J. & MANU, S. (2019) Morphological Characterization and Hydrodynamic Behavior of Shortfin Mako Shark (Isurus oxyrinchus) Dorsal Fin Denticles. Journal of Bionic Engineering, 16 (4): 730-741 http://dx.doi.org/10.1007/s42235-019-0059-7
ROWLEY, A. & LOCATELLO, L. & KAHRL, A. & REGO, M. & BOUSSARD, A. & GARZA-GISHOLT, E. & KEMPSTER, R.M. & COLLIN, S.P. & GIACOMELLO, E. & FOLLESA, M.C. & PORCU, C. & EVANS, J.P. & HAZIN, F. & GARCIA-GONZALEZ, F. & DALY-ENGEL, T. & MAZZOLDI, C. & FITZPATRICK, J.L. (2019) Sexual selection and the evolution of sperm morphology in sharks. Journal of Evolutionary Biology, in press http://dx.doi.org/10.1111/jeb.13501
RUTLEDGE, K.M. (2019) A New Guitarfish of the Genus Pseudobatos (Batoidea: Rhinobatidae) with Key to the Guitarfishes of the Gulf of California. Copeia, 107 (3): 451-463   http://dx.doi.org/10.1643/CI-18-166
SCHMID, K. & ANDRADE, M. & MACHADO, F. & ARAUJO, J. & CORREA, E. & GIARRIZZO, T. (2019) Morphological abnormality in a Longnose Stingray Hypanus guttatus (Bloch & Schneider, 1801) (Myliobatiformes: Dasyatidae). Biota Neotropica, 19 (4): e20190792 http://dx.doi.org/10.1590/1676-0611-bn-2019-0792
SCULLY, R.P. (2019) Sharks give off a ghostly green glow in deep waters. New Scientist, 243 (3243): 17-17 
SHAMIR, Z.Z. & SHAMIR, S.Z. & BECKER, N. & SCHEININ, A. & TCHERNOV, D. (2019) Evidence of the impacts of emerging shark tourism in the Mediterranean. Ocean & Coastal Management, 178: 104847 http://dx.doi.org/10.1016/j.ocecoaman.2019.104847
SHIPLEY, O.N. & GALLAGHER, A.J. & SHIFFMAN, D.S. & KAUFMAN, L. & HAMMERSCHLAG, N. (2019) Diverse resource-use strategies in a large-bodied marine predator guild: evidence from differential use of resource subsidies and intraspecific isotopic variation. Marine Ecology Progress Series, 623: 71-83 http://dx.doi.org/10.3354/meps12982
SIMPSON, S.J. & SIMS, D.W. & TRUEMAN, C.N. (2019) Ontogenetic trends in resource partitioning and trophic geography of sympatric skates (Rajidae) inferred from stable isotope composition across eye lenses. Marine Ecology Progress Series, 624: 103-116 http://dx.doi.org/10.3354/meps13030
SOARES, K.D.A. (2019) Sexually dimorphic body proportions in the catshark genus Scyliorhinus (Chondrichthyes: Carcharhiniformes: Scyliorhinidae). Journal of Fish Biology, 95 (2): 683-685 http://dx.doi.org/10.1111/jfb.14011
SPEED, C.W. & REES, M.J. & CURE, K. & VAUGHAN, B. & MEEKAN, M.G. (2019) Protection from illegal fishing and shark recovery restructures mesopredatory fish communities on a coral reef. Ecology and Evolution, in press http://dx.doi.org/10.1002/ece3.5575
TRUELOVE, N.K. & ANDRUSZKIEWICZ, E.A. & BLOCK, B.A. (2019) A rapid environmental DNA method for detecting white sharks in the open ocean. Methods in Ecology and Evolution, 10 (8): 1128-1135 http://dx.doi.org/10.1111/2041-210x.13201
TUCKER, J.P. & VERCOE, B. & SANTOS, I.R. & DUJMOVIC, M. & BUTCHER, P.A. (2019) Whale carcass scavenging by sharks. Global Ecology and Conservation, 19: e00655 http://dx.doi.org/10.1016/j.gecco.2019.e00655
VILLAFAÑA, J.A. & MARRAMÀ, G. & HERNANDEZ, S. & CARRILLO-BRICEÑO, J.D. & HOVESTADT, D. & KINDLIMANN, R. & KRIWET, J. (2019) The Neogene Fossil Record of Aetomylaeus (Elasmobranchii, Myliobatidae) from the Southeastern Pacific Journal of Vertebrate Paleontology, 39 (1): e1577251 http://dx.doi.org/10.1080/02724634.2019.1577251
WHITE, W.T. & KYNE, P.M. & HARRIS, M. (2019) The correct spelling of the nomen of the lost shark Carcharhinus obsolerus White, Kyne & Harris, 2019 (Chondrichthyes, Carcharhinidae). Zootaxa, 4657 (3): 599-600 http://dx.doi.org/10.11646/zootaxa.4657.3.14
WHITEHEAD, D.A. & RAMIREZ, D.P. & OLIVIER, D. & ARMAS, R.G. & PANCALDI, F. & GALVAN-MAGANA, F. (2019) Seasonal trends in whale shark Rhincodon typus sightings in an established tourism site in the Gulf of California, Mexico. Journal of Fish Biology, in press http://dx.doi.org/10.1111/jfb.14106
YEMISKEN, E. & NAVARRO, J. & FORERO, M. & MEGALOFONOU, P. & ERYILMAZ, L. (2019) Trophic partitioning between abundant demersal sharks coexisting in the North Aegean Sea. Journal of the Marine Biological Association of the United Kingdom, 99 (5): 1213-1219 http://dx.doi.org/10.1017/s0025315419000110
YOSEF, R. (2019) Tiger sharks eat songbirds: comment. Ecology, in press http://dx.doi.org/10.1002/ecy.2846

Extinct Chondrichthyes:
ENGELBRECHT, A. & MORS, T. & REGUERO, M.A. & KRIWET, J. (2019) Skates and rays (Elasmobranchii, Batomorphii) from the Eocene La Meseta and Submeseta formations, Seymour Island, Antarctica. Historical Biology, 31 (8): 1028-1044 http://dx.doi.org/10.1080/08912963.2017.1417403
FISCHER, J. & HARTKOPF-FRÖDER, C. & LEIPNER, A. & SCHNEIDER, J.W. & WITTRY, J. & SOWIAK, M. (2019) A highly diverse chondrichthyan egg capsule assemblage from the Late Pennsylvanian Piesberg Quarry, Northwest Germany. in: Kölner Forum Geol. Paläont., 23 (2019), S. HARTENFELS, H.-G. HERBIG, M.R.W. AMLER & M. ARETZ (Eds.), Abstracts, 19th International Congress on the Carboniferous and Permian, Cologne, July 29 – August 2, 2019  
MANGANELLI, G. & SPADINI, V. (2019) Megascyliorhinus miocaenicus (Chondrichthyes, Galeomorphii) from the Zanclean (early Pliocene) of San Quirico d'Orcia, central Italy. Bollettino Della Societa Paleontologica Italiana, 58 (2): 165-170  http://dx.doi.org/10.4435/bspi.2019.12
TAPANILA, L. & PRUITT, J. (2019) Redefining species concepts for the Pennsylvanian scissor tooth shark, Edestus. PLoS ONE, 14 (9): e0220958 http://dx.doi.org/10.1371/journal.pone.0220958
VILLAFAÑA, J.A. & NIELSEN, S.N. & KLUG, S. & KRIWET, J. (2019) Early Miocene cartilaginous fishes (Chondrichthyes: Holocephali, Elasmobranchii) from Chile: Diversity and palaeobiogeographic implications. Journal of South American Earth Sciences, in press http://dx.doi.org/10.1016/j.jsames.2019.102317


CARRASSON, M. & DALLARES, S. & CARTES, J.E. & CONSTENLA, M. & PEREZ-DEL-OLMO, A. & ZUCCA, L. & KOSTADINOVA, A. (2019) Drivers of parasite community structure in fishes of the continental shelf of the Western Mediterranean: the importance of host phylogeny and autecological traits. International Journal for Parasitology, 49 (9): 669-683 http://dx.doi.org/10.1016/j.ijpara.2019.04.004
SMALES, L.R. & BARTON, D.P. & CHISHOLM, L.A. (2019) Acanthocephalans from Australian elasmobranchs (Chondrichthyes) with a description of a new species in the genus Gorgorhynchus Chandler, 1934 (Rhadinorhynchidae). Systematic Parasitology, 96 (7): 565-573  http://dx.doi.org/10.1007/s11230-019-09871-x



Longline fishing hampering shark migration

Date: August 20, 2019
Source: University of Queensland
Summary: Longline fisheries around the world are significantly affecting migrating shark populations, according to an international study. The study found that approximately a quarter of the studied sharks' migratory paths fell under the footprint of longline fisheries, directly killing sharks and affecting their food supply.
full story

Great white sharks have suddenly disappeared from one of their favorite hangouts

Sightings of white sharks (Carcharodon carcharias) have crashed this year in False Bay near Cape Town, South Africa—one of the best-known hot spots of the predators in the world—and scientists aren’t sure why. Orcas, which love to dine on shark liver, may have scared them off, researchers say, but human activities could also play a role.

full story