Liu, F.-H. (1932)
The Elasmobranchiate fishes of north China. The Science reports of National Tsing Hua University. Series B: Biological and psychological sciences, 1(5): 133–177, Pl. 1–6.
Ishiyama, R. (1950)
Studies on the rays and skates belonging to the family Rajidae, found in Japan and adjacent regions. 1. Egg-capsules of ten species. Japanese Journal of Ichthyology, 1(1): 30–36
Ishiyama, R. (1958)
Studies on the rajid fishes (Rajidae) found in the waters around Japan. Journal of the Shimonoseki College of Fisheries, 7(2/3): 193–394, pls. 1–3
Ishiyama, R. (1958)
Observations on the egg-capsules of skates of the family Rajidae, found in Japan and its adjacent waters. Bulletin of the Museum of Comparative Zoology at Harvard College, 118(1): 1–24
Ishihara, H. & Ishiyama, R. (1986)
Systematics and distribution of the skates of the north Pacific. In: Uyeno, T. & Arai, R. & Taniuchi, T. & Matsuura, K.(eds) Proceedings of the second international conference on Indo-Pacific fishes. Ichthyological Society of Japan, Tokyo: 269–280
Amaoka, K. & Nakaya, K. & Yabe, M. (1989)
Fishes of Usujiri and Adjacent Waters in Southern Hokkaido, Japan. Bulletin of the Faculty of Fishery of the Hokkaido University, 40(4): 254–277
McEachran, J.D. & Konstantinou, H. (1996)
Survey of the variation of alar and malar thorns in skates: phylogenetic implications (Chondrichthyes: Rajoidei). Journal of Morphology, 228(2), 165–178
DOI: 10.1002/(SICI)1097-4687(199605)228:2<165::AID-JMOR5>3.0.CO;2-4
Mould, B. (1997)
Classification of the recent Elasmobranchii. Copyright Brian Mould 1997
Sokolovskaya, T.G. & Sokolovskii, A.S. & Sobolevskii, E.I. (1998)
A list of fishes of Peter the Great Bay (the Sea of Japan). Journal of Ichthyology, 38(1), 1–11
Dolganov, V.N. (1999)
Geographical and bathymetric distribution of the skates of the Rajidae family in the Far Eastern Seas of Russia and adjacent waters. Journal of Ichthyology, 39(4), 340–342
Deynat, P.P. (2000)
Les denticules myrmécoïdes, un nouveau caractère diagnostique pour les Rajidae (Chondrichtyes, Batoidea). Annales des Sciences Naturelles – Zoologie et Biologie Animale, 21(2): 65–80
DOI: 10.1016/S0003-4339(00)00106-4
Yoda, M. & Tokimura, M. & Horikawa, H. & Yamada, U. (2002)
A catalogue of fishes from the East China and Yellow seas with their local names. Seikai National Fisheries Research Institute, Fisheries Research Agency, Nagasaki, iii+42 pp.(in Japanese)
Compagno, L.J.V. (2005)
Checklist of living Chondrichthyes. In: Hamlett, W.C.(ed.) Reproductive biology and phylogeny of chondrichthyes: sharks, rays and chimaeras, Vol. 3. Endfield, USA: Science Publishers: 503–548
Shinohara, G. & Sato, T. & Aonuma, Y. & Horikawa, H. & Matsuura, K. & Nakabo, T. & Sato, K. (2005)
Annotated checklist of deep-sea fishes from the waters around the Ryukyu Islands, Japan. Deep-sea fauna and pollutants in the Nansei Islands. Monographs of the National Science Museum Tokyo, 29: 385–452
Ebert, D.A. & Bizzarro, J.J. (2007)
Standardized diet compositions and trophic levels of skates (Chondrichthyes: Rajiformes: Rajoidei). Environmental Biology of Fishes, 80(2–3), 221–237
DOI: 10.1007/s10641-007-9227-4
Ishihara, H. & Wang, Y. & Tanaka, Sho & Nakaya, K. & Jeong, C-H. (2009)
Raja pulchra In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.2. <www.iucnredlist.org>
Yoon, H.K. & Jeong, D. & Chung, I. & Jung, J.W. & Oh, M.J. & Kim, S. & Lee, Y.-H. & Kim, C.-G. & Hwang, S.Y. (2009)
Rapid Species Identification of Elasmobranch Fish (Skates and Rays) using Oligonucleotide Microarray. Biochip Journal, 3(1): 87–96
Antonenko, D.V. & Solomatov, S.V. & Balanov, A.A. & Kim Sen Tok & Kalchugin, P.V. (2011)
Occurrence of skate Raja pulchra (Rajidae, Rajiformes) in Russian waters of the Sea of Japan. Journal of Ichthyology, 51(6), 426–431
DOI: 10.1134/S0032945211040011
Chiquillo, K. & Crow, K. & Bravo, N. & Ebert, D.A. (2012)
The secret of the Mermaid's Purse: A novel reproductive strategy in two skates of the genus Raja. Abstract - Poster World Congress of Herpetology, Vancouver, Canada, August 8 – 14, 2012, Book of Abstracts: 131–132
Ishihara, H. & Treloar, M. & Bor, P.H.F. & Senou, H. & Jeong, C.-H. (2012)
The Comparative Morphology of Skate Egg Capsules (Chondrichthyes: Elasmobranchii: Rajiformes). Bulletin of the Kanagawa Prefectural Museum, Natural Science, 41, 9–25
Li, L. & Gibson, D.I. & Liu, Y.-Y. & Zhang, L.-P. (2012)
Morphological and molecular study of the poorly known species Pseudanisakis rajae (Yamaguti, 1941) (Nematoda: Acanthocheilidae) from elasmobranchs in the Yellow Sea and Taiwan Strait off the coast of China. Systematic Parasitology, 81(2), 115–123
DOI: 10.1007/s11230-011-9331-x
Grigorov, I.V. & Orlov, A.M. (2013)
Species diversity and conservation status of cartilaginous fishes (Chondrichthyes) of Russian waters. Journal of Ichthyology, 53(11), 923–936
DOI: 10.1134/S0032945213110040
Nakabo, T. (2013)
Fishes of Japan with Pictorial Keys to the Species. Third Edition. Tokyo: Tokai University Press. 2428 pp.
Chiquillo, K.L. & Ebert, D.A. & Slager, C.J. & Crow, K.D. (2014)
The secret of the mermaid’s purse: Phylogenetic affinities within the Rajidae and the evolution of a novel reproductive strategy in skates. Molecular Phylogenetics and Evolution, 75: 245–251
DOI: 10.1016/j.ympev.2014.01.012
Parin, N.V. & Evseenko, S.A. & Vasil'eva, E.D. (2014)
Fishes of Russian Seas: Annotated Catalogue. KMK Scientific Press, Moscow v. 53: 733 pp.
Dyldin, Y.V. (2015)
Annotated checklist of the sharks, batoids and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali) from waters of Russia and adjacent areas. Publications of the Seto Marine Biological Laboratory, 43: 40–91
Hwang, I.K. & Lee, H.Y. & Kim, M.-H. & Jo, H.-S. & Choi, D.-H. & Kang, P.-W. & Lee, Y.-H. & Cho, N.-S. & Park, K.-W. & Chae, H.Z. (2015)
Development of real-time PCR assay for genetic identification of the mottled skate, Beringraja pulchra. Forensic Science International, 255, 80–84
DOI: 10.1016/j.forsciint.2015.05.028
Im, Y.-J. & Jo, H.–S. (2015)
Migration and growth rate of Mottled skate, Beringraja pulchra by the tagging release program in the Yellow Sea, Korea. Bulletin of the Korean Society of Fisheries Technology, 51(2): 227–234
DOI: 10.3796/ksft.2015.51.2.227
Lim, Y.-J. & Jo, H.-S. & Jeong, G.-S. & Hwang, B.-K. & Kang, S.I. & Heu, M.S. & Kim, J.-S. (2015)
Sensory and Nutritional Characterizations of Mottled Skate Beringraja pulchra Caught off Ulleung Island, Korea. Korean Journal of Fisheries and Aquatic Sciences, 48(3): 275–283
DOI: 10.5657/KFAS.2015.0275
Panchenko, V.V. & Boiko, M.I. (2015)
On Discovery of the Mottled Skate Raja pulchra off the Coast of Primorye (Sea of Japan). Journal of Ichthyology, 55(5), 747–749
DOI: 10.1134/S0032945215050112
Tohkairin, A. & Hamatsu, T. & Yoshikawa, A. & Kai, Y. & Nakabo, T. (2015)
An illustrated and annotated checklist of fishes on Kitami-Yamato Bank, southern Sea of Okhotsk. Publications of the Seto Marine Biological Laboratory, 43: 1–29
Hwang, J.Y. & Jin, G.-D. & Park, J. & Kim, H. & Lee, C.-K. & Kwak, W. & Nam, B.-H. & An, C.M. & Park, J.Y. & Park, K.-H. & Huh, C.-S. & Kim, E.B. (2016)
Complete genome sequence and SNPs of Raja pulchra (Rajiformes, Rajidae) mitochondria. Mitochondrial DNA, 27(4), 2975–2977
DOI: 10.3109/19401736.2015.1060467
Jeong, D. & Kim, S. & Kim, C.-G. & Myoung, J.-G. & Lee, Y.-H. (2016)
Complete mitochondrial genome of the mottled skate: Raja pulchra (Rajiformes, Rajidae). Mitochondrial DNA, 27(3), 1823–1824
DOI: 10.3109/19401736.2014.971242
Last, P.R. & Séret, B. & Stehmann, M.F.W. & Weigmann, S. (2016)
Skates, Family Rajidae. In: Last, P.R., White, W.T., Carvalho, M.R. de, Séret, B., Stehmann, M.F.W & Naylor, G.J.P(Eds.) Rays of the World. CSIRO Publishing, Melbourne: 204–363
Last, P.R. & Weigmann, S. & Yang, L. (2016)
Changes to the nomenclature of the skates (Chondrichthyes: Rajiformes). In Rays of the World: Supplementary information(Last, P. R. & Yearsley, G. K., eds), CSIRO Special Publication: 11–34
Weigmann, S. (2016)
Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology, 88(3), 837–1037
DOI: 10.1111/jfb.12874
Amaral, C.R.L. & Pereira, F. & Silva, D.A. & Amorim, A. & de Carvalho, E.F. (2017)
The mitogenomic phylogeny of the Elasmobranchii (Chondrichthyes). Mitochondrial DNA Part A, 20(6), 1–12
DOI: 10.1080/24701394.2017.1376052
Im, Y.-J. & Jo, H.-S. & Ji, H.-S. & Myoung, S.-H. & Kim, J.-K. (2017)
Geographic variations of the mottled skate, Beringraja pulchra (Liu, 1932) (Rajidae) in the Yellow and East seas based on molecular and morphometric data. Journal of Applied Ichthyology, 33(5), 950–956
DOI: 10.1111/jai.13408
Jang, G.I. & Kim, G. & Hwang, C.Y. & Cho, B.C. (2017)
Prokaryotic community composition in alkaline-fermented skate (Raja pulchra). Food Microbiology, 61, 72–82
DOI: 10.1016/j.fm.2016.08.008
Amaral, C.R.L. & Pereira, F. & Silva, D.A. & Amorim, A. & Decarvalho, E.F. (2018)
The mitogenomic phylogeny of the Elasmobranchii (Chondrichthyes). Mitochondrial DNA Part A, 29(6), 867–878
DOI: 10.1080/24701394.2017.1376052
Barbini, S.A. & Sabadin, D.E. & Lucifora, L.O. (2018)
Comparative analysis of feeding habits and dietary niche breadth in skates: the importance of body size, snout length, and depth. Reviews in Fish Biology and Fisheries, 28(3), 625–636
DOI: 10.1007/s11160-018-9522-5
Buckley, K.A. & Crook, D.A. & Pillans, R.D. & Smith, L. & Kyne, P.M. (2018)
Sustainability of threatened species displayed in public aquaria, with a case study of Australian sharks and rays. Reviews in Fish Biology and Fisheries, 28(1), 137–151
DOI: 10.1007/s11160-017-9501-2
Dyldin, Y.V. & Orlov, A.M. (2018)
An Annotated List of Cartilaginous Fishes (Chondrichthyes: Elasmobranchii, Holocephali) of the Coastal Waters of Sakhalin Island and the Adjacent Southern Part of the Sea of Okhotsk. Journal of Ichthyology, 58(2), 158–180
Kim, M.R. & Kwon, K. & Jung, Y.K. & Kang, T.S. (2018)
A rapid real-time PCR method to differentiate between mottled skate (Beringraja pulchra) and other skate and ray species. Food Chemistry, 255, 112–119
DOI: 10.1016/j.foodchem.2018.02.056
Choi, S. & Kim, S.J. & Kim, E.B. (2019)
Complete mitochondrial DNA sequence of Raja pulchra from Yellow Sea and Alaska. Mitochondrial DNA Part B, 4(1), 384–385
DOI: 10.1080/23802359.2018.1547144
Li, W. & Kobayashi, T. & Moroi, S. & Kotake, H. & Ikoma, T. & Saeki, H. & Ura, K. & Takagi, Y. (2019)
Anti-obesity effects of chondroitin sulfate oligosaccharides from the skate Raja pulchra. Carbohydrate Polymers, 214: 303-310
DOI: 10.1016/j.carbpol.2019.03.025
Barbini, S.A. & Lucifora, L.O. & Sabadin, D.E. & Figueroa, D.E. (2020)
Ecological specialization is associated with high conservation concern in skates (Chondrichthyes, Rajiformes). Animal Conservation, 23(2), 222–228
DOI: 10.1111/acv.12531
Guinot, G. & Cavin, L. (2020)
Distinct Responses of Elasmobranchs and Ray-Finned Fishes to Long-Term Global Change. Frontiers in Ecololgy and Evolution, 7, Article 513
DOI: 10.3389/fevo.2019.00513
Park, J. & Kim, S.J. & Kim, E. (2020)
Changes in the Microbial Community of the Mottled Skate (Beringraja pulchra) during Alkaline Fermentation. Journal of Microbiology and Biotechnology, 30(8): 1195–1206
DOI: 10.4014/jmb.2003.03024
Sonoyama, T. & Ogimoto, K. & Hori, S. & Uchida, Y. & Kawano, M. (2020)
An annotated checklist of marine fishes of the Sea of Japan off Yamaguchi Prefecture, Japan, with 74 new records. (in Japanese) Bulletin of the Kagoshima University Museum, 11: 1-152