Shimada, K. & Motani, R. & Wood, J.J. & Sternes, P.C. & Tomita, T. & Bazzi, M. & Collareta, A. & Gayford, J.H. & Türtscher, J. & Jambura, P.L. & Kriwet, J. & Vullo, R. & Long, D.J. & Summers, A.P. & Maisey, J.G. & Underwood, C. & Ward, D.J. & Maisch, H.M. & Perez, V.J. & Feichtinger, I. & Naylor, G.J.P. & Moyer, J.K. & Higham, T.E. & da Silva, J.P.C.B. & Bornatowski, H. & González-Barba, G. & Griffiths, M.L. & Becker, M.A. & Siversson, M. (2025)
Reassessment of the possible size, form, weight, cruising speed, and growth parameters of the extinct megatooth shark, Otodus megalodon (Lamniformes: Otodontidae), and new evolutionary insights into its gigantism, life history strategies, ecology, and extinction Palaeontologia Electronica, 28(1), Article a12
DOI: 10.26879/1502

Tomita, T. & Yabumoto, Y. & Kuga, N. (2024)
A New Snaggletooth Shark Species, Hemipristis tanakai Sp. Nov., from the Ashiya Group (Oligocene), Northern Kyushu, Japan. Paleontological Research, 28(3), 273–278
DOI: 10.2517/PR220021

Sternes, P.C. & Jambura, P.L. & Türtscher, J. & Kriwet, J. & Siversson, M. & Feichtinger, I. & Naylor, G.J.P. & Summers, A.P. & Maisey, J.G. & Tomita, T. & Moyer, J.K. & Higham, T.E. & da Silva, J.P.C.B. & Bornatowski, H. & Long, D.J. & Perez, V.J. & Collareta, A. & Underwood, C. & Ward, D.J. & Vullo, R. & González-Barba, G. & Maisch, H.M. & Griffiths, M.L. & Becker, M.A. & Wood, J.J. & Shimada, K. (2024)
White shark comparison reveals a slender body for the extinct megatooth shark, Otodus megalodon (Lamniformes: Otodontidae). Palaeontologia Electronica, 27(1), Article a4
DOI: 10.26879/1345

Tomita, T. & Miyamoto, K. & Nakamura, M. & Murakumo, K. & Toda, M. & Sato, K. (2024)
Whitish film covering a newborn white shark was not intrauterine material but embryonic epithelium. Environmental Biology of Fishes, 107, 719–722
DOI: 10.1007/s10641-024-01560-z

Tomita, T. & Toda, M. & Kaneko, A. & Murakumo, K. & Miyamoto, K. & Sato, K. (2023)
Successful delivery of viviparous lantern shark from an artificial uterus and the self-production of lantern shark luciferin. Plos One, 18(9), Article e0291224
DOI: 10.1371/journal.pone.0291224

Tomita, T. & Nakamura, M. & Nozu, R. & Ogawa, N. & Toda, M. & Sato, K. (2022)
Mode of uterine milk secretion in the white shark. The Anatomical Record, 305(7), 1724–1731
DOI: 10.1002/ar.24860

Türtscher, J. & Jambura, P.L. & López-Romero, F.A. & Kindlimann, R. & Sato, K. & Tomita, T. & Kriwet, J. (2022)
Heterodonty and ontogenetic shift dynamics in the dentition of the tiger shark Galeocerdo cuvier (Chondrichthyes, Galeocerdidae). Journal of Anatomy, 241(2), 372–392
DOI: 10.1111/joa.13668

Tomita, T. & Toda, M. & Murakumo, K. & Kaneko, A. & Yano, N. & Nakamura, M. & Sato, K. (2022)
Five-Month Incubation of Viviparous Deep-Water Shark Embryos in Artificial Uterine Fluid. Frontiers in Marine Science, 9, Article 825354
DOI: 10.3389/fmars.2022.825354

Tomita, T. & Nakamura, M. & Miyamoto, K. & Nozu, R. & Toda, M. & Murakumo, K. & Sato, K. (2021)
Clasper pocket: adaptation of a novel morphological feature by lamnoid sharks, which aids with tuna-like swimming. Zoomorphology, 140, 365–371
DOI: 10.1007/s00435-021-00532-6

Tomita, T. & Toda, M. & Murakumo, K. & Miyamoto, K. & Matsumoto, R. & Ueda, K. & Sato, K. (2021)
Volume of the whale shark and their mechanism of vertical feeding. Zoology, 147, Article 125932
DOI: 10.1016/j.zool.2021.125932

Tomita, T. & Murakumo, K. (2021)
Where embryology meets functional morphology: A new frontier of science [in Japanese]. Report of Japanese Society for Elasmobranch Studies, 57, 2–6

Murakumo, K. & Matsumoto, R. & Tomita, T. & Matsumoto, Y. & Ueda, K. (2020)
The power of ultrasound: observation of nearly the entire gestation and embryonic developmental process of captive reef manta rays (Mobula alfredi). Fishery Bulletin, 118(1), 1–7
DOI: 10.7755/FB.118.1.1

Tomita, T. & Murakumo, K. & Komoto, S. & Dove, A. & Kino, M. & Miyamoto, K. & Toda, M. (2020)
Armored eyes of the whale shark. PLoS ONE, 15(6), Article e0235342
DOI: 10.1371/journal.pone.0235342

Tomita, T. & Nakamura, M. & Kobayashi, Y. & Yoshinaka, A. & Murakumo, K. (2020)
Viviparous stingrays avoid contamination of the embryonic environment through faecal accumulation mechanisms. Scientific Reports, 10, Article 7378
DOI: 10.1038/s41598-020-64271-2

Duchatelet, L. & Moris, V.C. & Tomita, T. & Mahillon, J. & Sato, K. & Behets, C. & Mallefet, J. (2020)
The megamouth shark, Megachasma pelagios, is not a luminous species. PLoS ONE, 15(11), Article e0242196
DOI: 10.1371/journal.pone.0242196

Tomita, T. & Murakumo, K. & Ueda, K. & Ashida, H. & Furuyama, R. (2019)
Locomotion is not a privilege after birth: Ultrasound images of viviparous shark embryos swimming from one uterus to the other. Ethology, 125(2), 122–126
DOI: 10.1111/eth.12828

Duchatelet, L. & Pinte, N. & Tomita, T. & Sato, K. & Mallefet, J. (2019)
Etmopteridae bioluminescence: dorsal pattern specificity and aposematic use. Zoological Letters, 5, Article 9
DOI: 10.1186/s40851-019-0126-2

Tomita, T. & Toda, M. & Miyamoto, K. & Oka, S.-I. & Ueda, K. & Sato, K. (2018)
Development of the Lunate-Shaped Caudal Fin in White Shark Embryos. The Anatomical Record, 301(6), 1068–1073
DOI: 10.1002/ar.23776

Tomita, T. & Toda, M. & Miyamoto, K. & Ueda, K. & Nakaya, K. (2018)
Morphology of a hidden tube: Resin injection and CT scanning reveal the three-dimensional structure of the spiracle in the Japanese bullhead shark Heterodontus japonicus (Chondrichthyes; Heterodontiformes; Heterodontidae). The Anatomical Record, 301(8), 1336–1341
DOI: 10.1002/ar.23836

Tomita, T. & Touma, H. & Murakumo, K. & Yanagisawa, M. & Yano, N. & Oka, S.-I. & Miyamoto, K. & Hanahara, N. & Sato, K. (2018)
Captive Birth of Tiger Shark (Galeocerdo cuvier) Reveals a Shift in Respiratory Mode during Parturition. Copeia, 106(2), 292–296
DOI: 10.1643/CI-17-683

Hirasaki, Y. & Tomita, T. & Yanagisawa, M. & Ueda, K. & Sato, K. & Okabe, M. (2018)
Heart Anatomy of Rhincodon typus: Three-Dimensional X-Ray Computed Tomography of Plastinated Specimens. The Anatomical Record, 301(11), 1801–1808
DOI: 10.1002/ar.23902

Tomita, T. & Toda, M. & Murakumo, K. (2018)
Stealth breathing of the angelshark. Zoology, 130, 1–5
DOI: 10.1016/j.zool.2018.07.003

Tomita, T. & Miyamoto, K. & Kawaguchi, A. & Toda, M. & Oka, S.-I. & Nozu, R. & Sato, K. (2017)
Dental ontogeny of a white shark embryo. Journal of Morphology, 278(2), 215–227
DOI: 10.1002/jmor.20630

Tomita, T. & Nozu, R. & Nakamura, M. & Matsuzaki, S. & Miyamoto, K. & Sato, K. (2017)
Live-bearing without placenta: Physical estimation indicates the high oxygen-supplying ability of white shark uterus to the embryo. Scientific Report, 7(1): 11744
DOI: 10.1038/s41598-017-11973-9

Tomita, T. & Cotton, C.F. & Toda, M. (2016)
Ultrasound and physical models shed light on the respiratory system of embryonic dogfishes. Zoology, 119(1), 36–41
DOI: 10.1016/j.zool.2015.09.002

Tomita, T. & Murakumo, K. & Miyamoto, K. & Sato, K. & Oka, S.I. & Kamisako, H. & Toda, M. (2016)
Eye retraction in the giant guitarfish, Rhynchobatus djiddensis (Elasmobranchii: Batoidea): a novel mechanism for eye protection in batoid fishes. Zoology, 119(1), 30–35
DOI: 10.1016/j.zool.2015.09.004

Sato, K. & Nakamura, M. & Tomita, T. & Toda, M. & Miyamoto, K. & Nozu, R. (2016)
How great white sharks nourish their embryos to a large size: evidence of lipid histotrophy in lamnoid shark reproduction. Biology Open, 5, 1211–1215
DOI: 10.1242/bio.017939

Nakaya, K. & Tomita, T. & Suda, K. & Sato, K. & Ogimoto, K. & Chappell, A. & Sato, T. & Takano, K. & Yuki, Y. (2016)
Slingshot feeding of the goblin shark Mitsukurina owstoni (Pisces: Lamniformes: Mitsukurinidae). Scientific Reports, 6, Article 27786
DOI: 10.1038/srep27786

Tomita, T. & Yokoyama, K. (2015)
The First Cenozoic Record of a Fossil Megamouth Shark (Lamniformes, Megachasmidae) from Asia. Paleontological Research, 19(3), 204–207
DOI: 10.2517/2015PR004

Tomita, T. (2015)
Pectoral Fin of the Paleozoic Shark, Cladoselache: New Reconstruction Based on a Near-Complete Specimen. Journal of Vertebrate Paleontology, 35(5), Article e973029
DOI: 10.1080/02724634.2015.973029

Tomita, T. & Kawai, T. & Matsubara, H. & Kobayashi, M. & Katakura, S. (2014)
Northernmost record of a whale shark Rhincodon typus from the Sea of Okhotsk. Journal of Fish Biology, 84(1), 243–246
DOI: 10.1111/jfb.12273

Tomita, T. & Tanaka, Sho & Sato, K. & Nakaya, K. (2014)
Pectoral Fin of the Megamouth Shark: Skeletal and Muscular Systems, Skin Histology, and Functional Morphology. PLoS ONE, 9(1), Article e8620
DOI: 10.1371/journal.pone.0086205

Tomita, T. & Nakamura, M. & Sato, K. & Takaoka, H. & Toda, M. & Kawauchi, J. & Nakaya, K. (2014)
Onset of Buccal Pumping in Catshark Embryos: How Breathing Develops in the Egg Capsule. PLoS ONE, 9(10), Article e109504

Tomita, T. & Toda, M. & Yamamoto, Y. & Sato, K. & Uchida, S. & Nakaya, K. (2013)
A novel pharyngeal expansion mechanism in the yellow-spotted fanray, Platyrhina tangi (Elasmobranchii: Batoidea), with special reference to the function of the fifth ceratobranchial cartilage in batoids. Zoomorphology, 132(3), 317–324
DOI: 10.1007/s00435-012-0185-9

Tomita, T. (2013)
Intermittent swimming lifestyle in hybodontid sharks, reconstructed on the basis of gill arch and caudal fin morphologies [Abstract]. In Schwarz, C. & Kriwet, J. (eds), 6th International Meeting on Mesozoic Fishes – Diversification and Diversity Patterns, Vienna, Austria August 4th–10th, 2013, 66

Tomita, T. & Kawai, T. & Matsubara, H. & Nagata, R. (2013)
Occurrence of the Chilean devil ray Mobula tarapacana (Elasmobranchii: Batoidea: Myliobatiformes) in the Sea of Okhotsk: first record from cold temperate waters. Journal of Fish Biology, 83(3), 695–698
DOI: 10.1111/jfb.12205

Tanaka, K. & Shiina, T. & Tomita, T. & Suzuki, S. & Hosomichi, K. & Sano, K. & Doi, H. & Kono, A. & Komiyama, T. & Inoko, H. & Kulski, J.K. & Tanaka, Sho (2013)
Evolutionary Relations of Hexanchiformes Deep-Sea Sharks Elucidated by Whole Mitochondrial Genome Sequences. BioMed Research International, 2013, Article ID 147064
DOI: 10.1155/2013/147064

Tomita, T. & Toda, M. & Ueda, K. & Uchida, S. & Nakaya, K. (2012)
Live-bearing manta ray, how the embryo acquires oxygen without placenta and umbilical cord. Biology Letters, 8(5), 721–724
DOI: 10.1098/rsbl.2012.0288

Tomita, T. & Sato, K. & Suda, K. & Kawauchi, J. & Nakaya, K. (2012)
Feeding of the megamouth shark (Pisces: Lamniformes: Megachasmidae) predicted by its hyoid arch: A biomechanical approach [Abstract]. - Poster World Congress of Herpetology, Vancouver, Canada, August 8 – 14, 2012, Book of Abstracts, 725–726

Tomita, T. & Kurihara, K. (2011)
First Record of a Large Lamniform Shark Cretodus semiplicatus in the Pacific Region, from the Mikasa Formation (Lower Cenomanian), Hokkaido, Japan. Paleontological Research, 15(3), 181–184
DOI: 10.2517/1342-8144-15.3.181

Tomita, T. & Sato, K. & Suda, K. & Kawauchi, J. & Nakaya, K. (2011)
Feeding of the megamouth shark (Pisces: Lamniformes: Megachasmidae) predicted by its hyoid arch: A biomechanical approach. Journal of Morphology, 272(5), 513–524
DOI: 10.1002/jmor.10905

Tomita, T. (2011)
Mouth-Size Estimation of a Primitive Lamniform Shark, Protolamna: Low Trophic Position in Lamniform Shark Origin. Paleontological Research, 15(2), 68–76
DOI: 10.2517/1342-8144-15.2.068

Tanaka, K. & Shiina, T. & Tomita, T. & Kinjo, S. & Hosomichi, K. & Sano, K. & Doi, H. & Kono, A. & Komiyama, T. & Inoko, H. & Kulski, J.K. & Tanaka, Sho (2011)
Phylogeny of whole mitochondrial genomes from Hexanchiform sharks [Abstract]. (Poster) In Programme Booklet of The 15th Annual Scientific Conference of the European Elasmobranch Association, Berlin, 29.10.–30.10.2011

Tomita, T. & Oji, T. (2010)
Habitat Reconstruction of Oligocene Elasmobranchs from Yamaga Formation, Ashiya Group, Western Japan. Paleontological Research, 14(1), 69–80
DOI: 10.2517/1342-8144-14.1.069

Tomita, T. (2009)
Reconstruction of fossil shark ventilation systems based on gill arch morphology and evolutionary implications [Abstract]. Journal of Vertebrate Paleontology, 29(Supplement to Number 3), 191A