Lloris, D. (1986)
Ictiofauna demersal y aspectos biogeográficos de la costa Sudoccidental de Africa (SWA/NAMIBIA). Monografías de zoología marina, Instituto de Ciencias del Mar, Barcelona, Consejo Superior de Investigaciones Científicas, 1: 429pp
Compagno, L.J.V. & Ebert, D.A. & Smale, M.J. (1989)
Guide to the sharks and rays of Southern Africa. New Holland Ltd., London 1989
Compagno, L.J.V. & Ebert, D.A. & Cowley, P.D. (1991)
Distribution of offshore demersal cartilaginous fishes (Class Chondrichthyes) of the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11(1), 43–139, figs 1–24
DOI: 10.2989/025776191784287664
Stehmann, M. (1995)
First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Archive of Fishery and Marine Research, 43(1), 1–119
McEachran, J.D. & Konstantinou, H. (1996)
Survey of the variation of alar and malar thorns in skates: phylogenetic implications (Chondrichthyes: Rajoidei). Journal of Morphology, 228(2), 165–178
DOI: 10.1002/(SICI)1097-4687(199605)228:2<165::AID-JMOR5>3.0.CO;2-4